The origin of this instinct, which helps to keep even closely allied
species in a state of nature distinct, seems to be sufficiently clear.
The number of species which have proved fertile together are very
limited, and the fertility of the hybrid offspring is almost constantly
diminished, often even to a very great extent. Of course, no one
now talks of the sterility of hybrids as a moral necessity—hybrids
being _animalia adulterina_,—or as the result of a special divine
decree, that new species should not be multiplied indefinitely.[1636]
M. Isidore Geoffroy has shown not only that hybrids may be fertile,
but that “infertile” hybrids are, properly speaking, merely the
hybrids which are most rarely fertile, their sterility never being
absolute.[1637] Moreover, as has been pointed out by Mr. Wallace, in
almost all the experiments that have hitherto been made in crossing
distinct species, no care has been taken to avoid close interbreeding;
hence these experiments cannot be held to prove that hybrids are
in all cases infertile _inter se_.[1638] But looking to all the
ascertained facts on the intercrossing of plants and animals, we may
with Mr. Darwin conclude that some degree of sterility in hybrids is an
extremely general result.[1639] This being the case with the hybrids of
our domesticated animals, it must be so all the more with animals in a
state of nature, which generally live under conditions less favourable
to mutual fertility. It is easy to understand, then, that instincts
leading to intercrossing of different species, even if appearing
occasionally, never could be long-lived, as only those animals which
preferred pairing with individuals of their own species, gave birth
to an offspring endowed with a normal power of reproduction, and thus
became the founders of numerous generations that inherited their
instincts.
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