The History of the European FaunaScharff, R. F. (Robert Francis)
History
The History of the European Fauna
Scharff, R. F. (Robert Francis)
Animals -- Europe; Zoogeography
Among the beetles belonging to this migration, there is one of very
considerable interest from a distributional point of view, for all the
species of the genus--even the whole family to which the genus
belongs--are what is known by zoologists as "Commensalists." These are
animals habitually associating and living in close connection with
others with which they are not tied by any family relations or kinship.
Such a state of close and permanent friendship is called "commensalism."
Now it appears as if the members of this family of beetles
(_Clavigeridæ_) had of their own free will formed such a close
connection with colonies of ants--sometimes with one species, sometimes
another. They are the permanent guests of the ants, and in return they
secrete a fluid which is apparently highly prized by them. All of the
_Clavigers_ are provided with peculiar club-shaped antennæ, with which
they ungraciously beat their hosts, when they are in want of food.
According to some authorities, they even occasionally gnaw at the pupæ
and larvæ of the ant with which they live.
Such beetles naturally can only have extremely limited means of
distribution, and they are comparable in that respect with the woodlice
of the genus _Platyarthrus_, to which I have already had occasion to
refer. All the species of _Claviger_ are confined to Europe, chiefly to
the south, but one species, _Cl. testaceus_, has wandered farther north
and occurs in the nest of the ant _Lasius flavus_ in the south of
England, Ireland, and Scotland. Though none of the _Clavigers_ can be
claimed as Oriental migrants, the centre of distribution of the genera
belonging to the _Clavigeridæ_ is in Southern Asia, and it is probable
that the ancestors of the European _Clavigers_ have spread westward from
that region to Europe, eastward to Australia and Japan, and southward to
Madagascar and South Africa. The genus _Hopatroides_, belonging to the
same family as the so-called Spanish-fly (_Tenebrionidæ_), has twelve
species in Western Asia and Greece. One only, _H. thoracicus_--an
instance of discontinuous distribution--occurs in Andalusia. _Amphicoma_
is represented in Western Asia and the Balkan peninsula by fifteen
species, while three others are met with in North-west Africa and
Southern Spain.
A genus of Dragon-fly, _Onychogomphus_, has in Europe a somewhat similar
distribution to _Claviger_, but it has besides a very extensive foreign
range. There are altogether thirty-five species; of these ten are
Holarctic, twelve Oriental, five Mascarene, and eight Ethiopian. The
centre of distribution is therefore in the Oriental region, and we may
assume that in all probability the genus has originated there, the
European species having travelled west with the Oriental migration at an
early date of the Tertiary Era.
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