The honey-bee: its nature, homes and productsHarris, William Hetherington
Science
The honey-bee: its nature, homes and products
Harris, William Hetherington
Bee culture; Bees
Passing next to cases in which the stamens and pistils of the
same flower come to maturity at different times, so as to make
cross-breeding a necessity, we may mention first some plants in which
the pollen ripens before the stigma is ready to receive it. We have
such a condition of things in the _willow-herb_, or _epilobium tribe_.
The pretty pink blossoms of a large variety of this genus are to be
found, in summer, along the banks of brooks and running ditches. We
will confine our remarks to the species distinguished by its narrow
leaves, and hence named _angustifolium_. When the flower has fully
opened, the eight stamens spread out, and their anthers shed the
pollen. Bees visit the blossoms, and getting dusted with the grains,
carry these away to other flowers of the same kind. And here, in
passing, we may recall the fact of bees keeping to one species of plant
during the whole of any one journey from the hive. The importance
of this can be now better appreciated, when its influence on the
fructification of blossoms is observed. But to return, the pistil of
the willow-herb remains, till the stamens have withered, curved round
out of the way, and unable to receive any of their pollen. Then, after
they are dead, it comes into such a position that it can take what
pollen may be brought to it from younger flowers. For the conveyance of
this it is dependent chiefly on bees, who do not fail to carry enough
for the required purpose. In this way each blossom, by the agency of
these insects, both gives and gets what is necessary for the continued
life of the species; and without these unconsciously conferred benefits
from insect life, no seeds of this kind of _epilobium_ would mature.
Another instance, equally interesting, is seen in the well-known
_cineraria_ tribe. The plants of this genus belong to the _composite_
order, in which what is usually called the blossom, consists of many
flowers, grouped together on one head. In the example before us, there
are nearly 200 thus aggregated. These florets separately open at
different times, those of the outer circles coming before those nearer
the centre. The pollen-tube of each is formed by five anthers, fastened
together at their edges, and discharging their pollen into the space
between them. At the lower part of this inclosure the pistil is
growing, but is not in a condition to receive usefully the fertilising
powder. It, however, as it advances, sweeps out, and carries up with
it, the pollen-grains, so that they may be conveyed to other florets of
the same or other blossoms, to effect their vitalising work. At length
the pistil, with its brush on its summit, comes into view, but, even
yet, is not sufficiently developed for fertilisation. In due course,
however, the upper end splits, and exposes the surface of the stigma
ready for the pollen, which must be brought from some other floret, and
probably from some other blossom. Thus cross-breeding is effectually
secured.
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