Setting aside the marsupials and edentata, to which we shall return,
it is true that the great normal orders of terrestrial mammals,
the ruminants, rodents, insectivora, and carnivora are almost
as cosmopolitan as man. But this is no longer the case with the
cheiroptera, not one of which passes the polar circle. As to the
quadrumana, it is well-known that they are wanting in Europe, with
the exception of the Rock of Gibraltar, in North America and in the
greater part of Asia and Oceania. Thus it appears that, even in the
extreme hypothesis which I have here indicated, it would not be in
the animal types which present the greatest resemblance to man, but
among the carnivora or ruminants that we should be forced to seek for
geographical analogies in favour of the pretended cosmopolitanism of
the _human order_.
V. This limitation of the areas of habitation of animals, which
is evidently related to their degree of elevation in the scale of
beings, is a general fact which we also meet with in plants. On this
point Ad. de Candolle speaks as follows:—“The mean area of species
is smaller according as the class to which they belong has a more
complete, a more highly developed, or, in other words, a more perfect
organisation.”
The _progressive localisation_ of organised beings, increasing
in degree as they become more perfect, is, then, a general law.
Physiology will readily account for this fact.
The perfection of organisms is the result of division of labour,
which demands the multiplication of functional apparatus. As the
anatomical instruments become more numerous and special, the
functions do the same. From this cause alone the conditions of
harmony between the living being and the conditions of life which
surround it become more and more definite. Consequently, the animal
or the plant only finds its really favourable conditions in a
constantly diminishing area. Beyond these limits the conditions of
life change, the struggle for existence becomes more hazardous, and
the spread of the species, genus, family, or even order is arrested.
Man alone, armed against the conditions of life by his _intelligence_
and _industry_, is capable of overcoming conditions of existence
which would be an impassable barrier _to his material organisation_.
The law of progressive localisation is in direct opposition to the
doctrine of the original cosmopolitanism of the human species.
In putting it aside, polygenists, properly so called, might draw
attention to the diffusion of the genera of dolphins and rorquals;
polygenistic monogenists of the school of Agassiz might argue from
the facts mentioned above in connection with the genera of megaptera
and sibaldius; they might both say: The general law of localisation
offers two exceptions; why should not man form a third?
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