Fertility, and the number of succeeding generations is increased,
when a superiority is given to one of the crossed species over the
other. This fact has been recognised in plants, and we meet with
it again in animals. By crossing and recrossing in a fixed manner
the goat and the sheep, hybrids called _chabins_ are obtained which
possess three-eighths of the paternal and five-eighths of the
maternal blood. These animals produce a fleece much valued in South
America, and are the source of real industry. They can be maintained
for several generations, but at length all the crossings to which
they owe their existence must be recommenced, they having returned to
the parental types, ‘like plants,’ as M. Gay said.
This proportion—three-eighths to five-eighths—appears to be very
favourable to the maintenance of hybrid races; it is the proportion
which characterises the famous leporides, the result of the crossing
of the hare and the rabbit. But can these hybrids, of which so
much has been said, maintain themselves without reverting to the
parental types? M. Roux evidently believed it, and it is still
asserted by M. Gayot. But the testimony of those who have established
and impugned their assertions leaves scarcely any room for doubt.
Isidore Geoffroy, who had at first believed in their fixity, and
had spoken of it as a conquest, did not hesitate afterwards to
admit the reversion. The fact has been established in the Jardin
d’Acclimatation, and M. Roux himself, upon the assertion of M.
Faivre, appears to have abandoned his previous assertions. The
observations and experiments made by the Agricultural Society of
Paris clearly show that the _leporides_, sent or presented by the
breeders themselves, had entirely reverted to the rabbit type.
Lastly, M. Sanson, discussing the anatomical side of the question,
has arrived at the same conclusions. Moreover, whoever will credit
the observations made by M. Naudin upon the Linariæ, will easily
recognise the _reversion_ and the _disordered variation_ exhibited
by the leporides of the Abbé Cagliari, who was the first to obtain a
fertile crossing between the hare and the rabbit.
These phenomena appear in an equally well marked manner in the
result of the cross between the silkmoth (_Bombyx cynthia_) and
the castor-oil silkmoth (_Bombyx arrindia_), obtained by M.
Guérin Méneville. The hybrids of the first generation were almost
exactly intermediate between the two species, and resembled each
other. In the second this uniformity disappeared, in the third the
dissimilarity had increased, some of the insects having reassumed
all the characters of the paternal or maternal types. In the seventh
generation this curious experiment was destroyed by ichneumons. But,
as M. Valée, their intelligent breeder, told me, nearly all the moths
had returned to the type of the Bombyx arrindia. The resemblance to
what took place in the case of M. Naudin’s Linariæ is here complete.
Public-domain text, read in full here on John Shaqi.
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