V. The phenomenon of the _reversion_ of the descendants of a hybrid
to the paternal or maternal type, or _disordered variation_, has
given rise to some interpretations which it will be well to rectify,
and has also raised important questions.
The attempt has been made to assimilate the latter to the
_oscillations_ presented by mongrels for some generations. But
daily experience should suffice to refute this opinion. Breeders
are crossing _races_ every day for some purpose or other, and they
would never do so if the crossing were to result in the production
of a disorder which would exhibit the smallest resemblance to that
displayed by the Linariæ of M. Naudin, and the silkmoths of Guérin
Méneville. They expect, however, a few irregularities more or less
marked, in the first generations, but they know that the race will
soon _settle_ while the disorder would only increase if the crossing
had taken place between _species_.
Again, an attempt has been made to consider the facts of _atavism_
and _reversion_ as identical. There is, however, a fundamental
difference between them, for the mongrel which by atavism reassumes
the characters of one of its paternal ancestors, for example, still
preserves its mixed nature. This is proved by the possibility of
its offspring of the first or second generation reproducing, on the
contrary, the essential traits of its own maternal ancestors. Darwin
gives many examples of facts of this nature from the agricultural
history of his country. One of the best to quote is that furnished by
the genealogy of a family of dogs observed by Girou de Buzareingues.
These animals were crosses between the setter and spaniel. Now one
male, a setter to all appearances, united with a female of pure
setter breed, produced spaniels, which makes it evident that the
latter blood was by no means annihilated, and that the return to the
setter type was only apparent.
It is different in the cases of _reversion_ displayed by hybrids,
for one of the two bloods is irrevocably expelled. We are justified
in making this assertion in the case of mammalia, by experience
extending as far back as the Roman period, or at least as far as the
seventeenth century. Titires and musmons have never since those times
had _offspring affected by atavism_. A ram and sheep have never been
known to produce a kid, nor a male and female goat to produce a lamb.
It is the same with plants, according to statements with which M.
Naudin has kindly furnished me.
Far from being similar, the phenomena of _atavism_ and _reversion_
are absolutely different and characteristic, the one of crossing
between _races_, the other of crossing between _species_. The first
proclaims the persistency of the physiological connections between
all the representatives, more or less modified, of one species; the
second proves the complete rupture of the same connections between
the descendants of two species accidentally brought into contact by
the promoter of the hybridism.
Public-domain text, read in full here on John Shaqi.
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