We know that with the exception of the marsupials (kangaroos,
sarrigue), and the monotremata (ornithorhynchus, echidna), all
mammals have a _placenta_, an organ essentially composed of a network
of blood-vessels, which unites the mother to the fœtus, and serves
for the nutrition of the latter. With the ruminants, the edentata,
and the cetacea, the placenta is _simple_ and _diffuse_, that is to
say, the tufts of the blood-vessels are developed upon the entire
surface of the fœtal envelope, and are in direct communication with
the inner surface of the uterus. In the rest of the mammals the
placenta is _double_; half being derived from the mother, and half
from the fœtus, or rather its external envelope. A special membrane
called the _Decidua_ covers the interior of the uterus, and unites
the placentæ. Haeckel, correctly attaching great importance to these
anatomical differences, divides mammals into two great groups: the
_indeciduata_, which have no decidua, and the _deciduata_, which
possess it.
Among the latter the placenta can surround the mammalian ovum like a
girdle (_zonoplacentalia_), or form a kind of circular disc more or
less developed (_discoplacentalia_). Man, apes, bats, insectivora,
and rodents, present the latter arrangement, and thus form a natural
group to which no _zonoplacential_, and, of course, no _indeciduate_
mammals can be admitted.
Haeckel, without the least hesitation, adds his _prosimiæ_ to the
groups which I have just enumerated, that is to say, he attributes
to them a decidua and a discoidal placenta. Now the anatomical
investigations of MM. Alphonse Milne Edwards and Grandidier upon the
animals brought by the latter from Madagascar place it beyond all
doubt that the prosimiæ of Haeckel have no decidua and a diffuse
placenta. They are _indeciduata_. Far from any possibility of their
being the ancestors of the apes, according to the principle laid down
by Haeckel himself, they cannot even be regarded as the ancestors of
the zonoplacential mammals, the carnivora for instance, and ought to
be connected with the pachydermata, the edentata and the cetacea.
Darwin and Haeckel will, perhaps reply that when they made their
genealogies, the embryogenesis of the prosimiæ was not known. But
why then represent them as one of the intermediate links to which
they attach so much importance? Their process is always the same,
considering the _unknown_ as a _proof_ in favour of their theory.
VI. The necessity, which I think has been clearly proved, of seeking
elsewhere than among the prosimiæ for the link which is required
between the marsupials and the apes, would not invalidate the
relationship between the latter and man. There are, however, other
facts which are irreconcilable with the theory.
Public-domain text, read in full here on John Shaqi.
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