The Life of CrustaceaCalman, W. T. (William Thomas)
Science
The Life of Crustacea
Calman, W. T. (William Thomas)
Crustacea
Belonging to a different category from any of those mentioned are
certain Crustacea closely allied to, or identical with, species living
in the sea, which inhabit inland lakes where no direct passage from the
sea is now possible. Attention was first called to these in the case of
some of the large lakes of Sweden, in which Professor Lovén found some
Crustacea--_Mysis relicta_ (see Fig. 16, p. 47), _Mesidotea entomon,
Pontoporeia affinis_--almost or quite identical with species inhabiting
the Baltic, the Arctic Ocean, and the North Atlantic. There is
geological evidence to show that these lakes were once fjords, or arms
of the sea, and have become cut off from communication with the Baltic
by gradual elevation of the land. The marine animals which they
contained would thus be imprisoned, and as the water became less and
less salt, by the inflow of rivers, certain species which were able to
accommodate themselves to the altered conditions would survive. Some of
the species living in the Swedish lakes have since been found to have a
wider distribution. Thus, _Mysis relicta_, which should perhaps be
reckoned as only a variety of the _Mysis oculata_ of Arctic seas, has
been found in lakes in Russia, North Germany, and North America (Lake
Superior and others), and has lately been discovered in Lough Neagh and
some other lakes in Ireland.
The brackish waters of the Caspian Sea contain a very remarkable
assemblage of animals, including many Crustacea, which, although now
quite isolated from the oceans, are certainly of marine, and in part of
Arctic, origin. Among these are some species closely allied to or
identical with those of the Swedish lakes already mentioned, together
with a great variety of species of Mysidacea, Cumacea, and Amphipoda,
which appear to have been evolved from marine forms since the Caspian
was cut off from communication with the Arctic Ocean.
To such assemblages of animals derived from marine species and isolated
in inland lakes the name of "relict" faunas has been given. It is
necessary to use caution, however, in extending this explanation of
their origin to every case of peculiar lake faunas. For example, there
are difficulties in the way of supposing that Lake Baikal was ever in
open and direct communication with the sea, although it contains many
animals, such as seals, which are certainly of marine origin. The chief
Crustacea of the lake are numerous species of Amphipods belonging to the
genus _Gammarus_, and other genera closely related thereto, and for
these, at all events, there is no need to assume a "relict" origin.
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