“Let us take any species,” he writes, on page 329 of _The Descent of Man_
(Ed. 1901), “a bird for instance, and divide the females inhabiting a
district into two equal bodies, the one consisting of the more vigorous
and better-nourished individuals, and the other of the less vigorous and
healthy. The former, there can be little doubt, would be ready to breed
in the spring before the others; and this is the opinion of Mr Jenner
Weir, who has carefully attended to the habits of birds during many
years. There can also be no doubt that the most vigorous, best nourished,
and earliest breeders would on an average succeed in rearing the largest
number of fine offspring. The males, as we have seen, are generally ready
to breed before the females; the strongest, and with some species the
best armed of the males, drive away the weaker; and the former would then
unite with the more vigorous and better-nourished females, because they
are the first to breed. Such vigorous pairs would surely rear a larger
number of offspring than the retarded females, which would be compelled
to unite with the conquered and less powerful males, supposing the sexes
to be numerically equal; and this is all that is wanted to add, in the
course of successive generations, to the size, strength, and courage of
the males, or to improve their weapons.”
From this competition among the males there arise, firstly, contests
between the males for mates; secondly, the preference of the females for
favoured males.
It is a matter of common knowledge that at the breeding season the males
of nearly all, if not all, species are very pugnacious. Two males often
engage in desperate fights for one or more females; the victor drives
away his foe and secures the harem. In such contests the stronger male
wins, and thus emerges that particular form of sexual selection which
Darwin termed “the law of battle.”
“There are,” writes Darwin, on page 324 of _The Descent of Man_, “many
other structures and instincts which must have developed through sexual
selection—such as the weapons of offence and the means of defence of the
males for fighting with and driving away their rivals—their courage and
pugnacity—their various ornaments—their contrivances for producing vocal
or instrumental music—and their glands for emitting odours.” The former
characters have, according to Darwin, been developed by the law of
battle, and the latter, since they serve only to allure or excite the
female, by the preference of the female.
Public-domain text, read in full here on John Shaqi.
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