“We can clearly discern this in the case of animals with simple habits.
Take the case of a carnivorous quadruped, of which the number that can be
supported in any country has long ago arrived at its full average. If its
natural power of increase be allowed to act, it can succeed in increasing
(the country not undergoing any change in its conditions) only by its
varying descendants seizing on places at present occupied by other
animals: some of them, for instance, being enabled to feed on new kinds
of prey, either dead or alive; some inhabiting new stations, climbing
trees, frequenting water, and some perhaps becoming less carnivorous. The
more diversified in habits and structure the descendants of our
carnivorous animal become, the more places they will be enabled to
occupy. What applies to one animal will apply throughout all time to all
animals—that is, if they vary—for otherwise natural selection can effect
nothing.” Darwin was, therefore, of opinion that natural selection is
able to bring about polytypic evolution. Darwin tacitly assumes, in the
illustration he gives, that the various races of the carnivorous animal
are in some way prevented from intercrossing; for if they interbreed
indiscriminately, these races will tend to be obliterated.
Isolation
“That perfectly free intercrossing,” writes Professor Lloyd Morgan (on p.
98 of _Animal Life and Intelligence_), “between any or all of the
individuals of a given group of animals is, so long as the characters of
the parents are blended in the offspring, fatal to divergence of
character, is undeniable. Through the elimination of less favourable
variations, the swiftness, strength, and cunning of a race may be
gradually improved. But no form of elimination can possibly differentiate
the group into swift, strong, and cunning varieties, distinct from each
other, so long as all three varieties freely interbreed, and the
characters of the parents blend with the offspring. Elimination may and
does give rise to progress in any given group, _as a group_; it does not
and cannot give rise to differentiation and divergence, so long as
interbreeding with consequent interblending of characters be freely
permitted. Whence it inevitably follows, as a matter of simple logic,
that where divergence has occurred, intercrossing and interbreeding must
in some way have been lessened or prevented.
“Thus a new factor is introduced, that of _isolation_ or _segregation_.
And there is no questioning the fact that it is of great importance. Its
importance, indeed, can only be denied by denying the swamping effects of
intercrossing, and such denial implies the tacit assumption that
interbreeding and interblending are held in check by some form of
segregation. The isolation explicitly denied is implicitly assumed.”
Public-domain text, read in full here on John Shaqi.
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