“The Indian robins present even greater difficulties to those who profess
to pin their faith to the all-sufficiency of natural selection. Robins
are found in nearly all parts of India, and fall into two species, the
brown-backed (_Thamnobia cambaiensis_) and the black-backed Indian Robin
(_Thamnobia fulicata_). The former occurs only in Northern India, and the
latter is confined to the southern portion of the peninsula. The hen of
each species is a sandy brown bird with a patch of brick-red feathers
under the tail, so that we cannot tell by merely looking at a hen to
which of the two species she belongs. The cock of the South Indian form
is, in winter, a glossy black bird, with a white bar in the wing, and the
characteristic red patch under the tail. The cock of the northern
species, as his name implies, has a sandy-brown back, which contrasts
strongly with the glossy black of his head, neck, and under parts. In
summer the cocks of the two species grow more like one another owing to
the wearing away of the outer edges of their feathers; but it is always
possible to distinguish between them at a glance. The two species meet at
about the latitude of Bombay. Oates states that in a certain zone, from
Ahmednagar to the mouth of the Godaveri valley, both species occur, and
they do not appear to interbreed.
“It seems impossible to maintain that natural selection, acting on minute
variations, has brought about the divergence between these two species.
Even if it be asserted that the difference in the colour of the feathers
of the back of the two cocks is in some way correlated with adaptability
to their particular environment, how are we to explain the fact that in a
certain zone both species flourish?
“A similar phenomenon is furnished by the red-vented bulbul. This genus
falls into several species, each corresponding to a definite locality and
differing only in details from the allied species, as, for example, the
distance down the neck to which the black of the head extends. There is a
Punjab Red-vented Bulbul (_Molpastes intermedius_), a Bengal (_Molpastes
bengalensis_), a Burmese (_Molpastes burmanicus_) and a Madras
(_Molpastes hæmorrhous_) species.
“It does not seem possible to maintain the contention that these various
species are the products of natural selection, for that would mean if the
black of the head of the Punjab species extended further into the neck
the bird could not live in that country.”
Thus, natural selection clearly is unable to explain some cases of
divergence of character due to geographical isolation.
There remains the third explanation, that the divergence is the result of
the simple fact of isolation.
We have already shown how insuperable are the objections to the view held
by Romanes and Gulick.
Public-domain text, read in full here on John Shaqi.
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