Seen under the magnifying-glass, it is a round, stiff, horny thread,
midway in thickness between a human hair and a horse-hair. Its tip is a
little rough, pointed and bevelled to some length down. The microscope
becomes necessary if we would see its real structure, which is much less
simple than it at first appears. We perceive that the bevelled end-part
consists of a series of truncated cones, fitting one into the other,
with their wide base slightly projecting. This arrangement produces a
sort of file, a sort of rasp with very much blunted teeth. When pressed
on the slide, the thread divides into four pieces of unequal length. The
two longer end in the toothed bevel. They come together in a very narrow
groove, which receives the two other, rather shorter pieces. These both
end in a point, which, however, is not toothed and does not project as
far as the final rasp. They also unite to form a groove, which fits into
the groove of the other two, the whole constituting a complete channel
or duct. Moreover, the two shorter pieces, considered together, can
move, lengthwise, in the groove that receives them; they can also move
one over the other, always lengthwise, so much so that, on the slide of
the microscope, their terminal points are seldom situated on the same
level.
If with our scissors we cut a piece of the inoculating-thread from the
living insect and examine the section under the magnifying-glass, we
shall see the inner groove lengthen out and project beyond the outer
groove and then go in again in turn, while from the wound there oozes
a tiny albimunous drop, doubtless proceeding from the liquid that gives
the egg the singular appendage to which we shall come presently. By
means of these longitudinal movements of the inner trench inside the
outer trench and of the sliding, one over the other, of the two portions
of the former, the egg can be despatched to the end of the ovipositor
notwithstanding the absence of any muscular contraction, which is
impossible in a horny conduit.
We have only to press the upper surface of the abdomen to see it
disjoint itself from the first segment, as though the insect had been
cut almost in two at that point. A wide gap or hiatus appears between
the first and second rings; and, under a thin membrane, the base of the
ovipositor bulges out, bent back into a stout hook. Here the filament
passes through the insect from end to end and emerges underneath. Its
issue is therefore near the base of the abdomen, instead of at the tip,
as usual. This curious arrangement has the effect of shortening the
lever-arm of the ovipositor and bringing the starting-point of the
filament nearer to the fulcrum, namely, the legs of the insect, and of
thus assisting the difficult task of inoculation by making the most of
the effort expended.
Public-domain text, read in full here on John Shaqi.
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