An osmotic growth also shows indications of movements which are connected
with its nutrition, and these movements are rhythmic, like those of
respiration or circulation in a living organism. The growth of an osmotic
production shows itself not as a continuous process but periodically. The
water traverses the membrane, raises the pressure, and distends the cell;
at first the cell wall resists by reason of its elasticity, it then
suddenly relaxes, yielding to the osmotic pressure and bulging out at a
thinner spot on the surface; the internal pressure falls suddenly, and
there is a pause in the growth.
This rhythmic growth may be best observed by sowing in a solution of a
tribasic alkaline phosphate, pellets composed of powdered calcium chloride
moistened with glycerine, to which has been added 1 per cent. of monobasic
calcium phosphate. The experiment is so arranged as to bend or incline the
growing stems which shoot out from these grains. This may be done by
carefully pouring above the mother liquor a layer of water, or a less
concentrated solution. As the internal osmotic pressure rises, the drooping
extremity of the twig will become turgescent and gradually lift itself
{157} up, and then suddenly fall again for several millimetres. We have
frequently watched this rhythmic movement for an hour or more--a slow
gradual elevation of the extremity of the twig and a rapid fall recurring
every four seconds or so.
It may be objected that the substance of an osmotic growth is continually
undergoing change, whereas a living organism transforms into its own
substance the extraneous matter which it borrows from its environment. The
distinction, however, is only an apparent one. The substance of a living
being is also continually undergoing chemical change; it does not remain
the same for a single instant. We see an evidence of this change in the
evolution of age; the substance of the adult is not that of the infant. In
some living organisms such as insects, especially the ephemeridae who have
but a brief existence, this change of substance is even more rapid than
that in an osmotic growth.
It has been objected that osmotic productions cannot be compared with
living organisms since they contain no albuminoid matter. This is to
consider life as a substance, and to confound the synthesis of life with
that of albumin. If albumin is ever produced by synthesis in the laboratory
it will probably be dead albumin. All living organisms contain albumin;
this is probably due to the fact that albuminoid matter is particularly
adapted for the formation of osmotic membranes. Our osmotic productions are
composed of the same elements as those which constitute living beings; an
osmotic growth obtained by sowing calcium nitrate in a solution of
potassium carbonate with sodium phosphate and sulphate contains all the
principal elements of a living organism, viz. carbon, oxygen, hydrogen,
nitrogen, sulphur, and phosphorus.
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