There is every reason to believe that the production of species is a slow process, and if fresh-water areas have not continued as a rule through long geological periods, we can see how variation has been constantly checked by the destruction, first in one part, then in another, of all the fresh-water species; and on these places being again occupied by fresh water they would be colonised by forms from other parts of the world. Thus species of restricted range were always exposed to destruction because their habitat was temporary and their retreat impossible, and only families of wide distribution could be preserved. Hence I believe it is that the types of fresh-water productions are few and world-wide, whilst the sea has mollusks innumerable, and the land great variety and wealth of species. This variety is in the ratio of the continuity of their habitats in time and space. It follows also, from the same reasoning, that old and widespread types are more likely to be preserved in fresh-water areas than on land or in the sea, for the destruction of wide-ranging species is effected more by the competition of improved varieties than by physical causes; so that when variation is most checked old forms will longest survive. Therefore I think it is that amongst fishes we find some old geological types still preserved in a few of the large rivers of the world. To illustrate more clearly the theory I have advanced, I will take a supposititious case. In the southern states of America there is reason to suppose that since the glacial period there has been a great variation in the species of the fresh-water mollusk genus Melania, and in different rivers there are distinct groups of species. Now let us suppose that the glacial period were to return, and that the icy covering, gradually thickening in the north, should push down southward as it did once before. The great lakes of North America would be again filled with ice, and their inhabitants destroyed. As the ice advanced southward, the inhabitants of one river-system after another would be annihilated, and many groups of Melania entirely destroyed. On the retreat of the ice again the rivers and lakes would reappear, but the varieties of animals that had been developed in them would not, and their places would be taken by aquatic forms from other areas, so that the number of species would be thereby greatly reduced, and wide-spreading forms would be freed from the competition of many improved varieties. Viewed in this light, the similarity of fresh-water productions all over the world, instead of being a difficulty in the way of the acceptance of the theory of natural selection, becomes a strong argument in favour of its truth; for we perceive that the number of marine, terrestrial, and freshwater animals is in proportion to the more or less continuous development that was possible under the different conditions under which they lived.