Agassiz, in his system of classification, has divided fishes into
four orders, according to the form of their scales; and his principle
of division, though apparently arbitrary and trivial, is yet found
to separate the class into great natural families, distinguished
from one another by other and very striking peculiarities. One kind
of scale, for instance, the placoid or broad-plated scale, is found
to characterize all the cartilaginous fishes of Cuvier except the
sturgeon;--it is the characteristic of an otherwise well-marked
series, whose families are furnished with skeletons composed of mere
animal matter, and whose gills open to the water by spiracles. The
fish of another order are covered by ctenoid or comb-shaped scales,
the posterior margin of each scale being toothed somewhat like the
edge of a saw or comb; and the order, thus distinguished, is found
wonderfully to agree with an order formed previously on another
principle of classification, the Acanthopterygii, or thorny-finned
order of Cuvier, excluding only the smooth-scaled families of
this previously formed division, and including, in addition to
it, the flat fish. A third order, the Cycloidean, is marked by
simple marginated scales, like those of the cod, haddock, whiting,
herring, salmon, &c.; and this order is found to embrace chiefly
the Malacopterygii, or soft-finned order of Cuvier--an order to
which all these well-known fish, with an immense multitude of others,
belong. Thus the results of the principle of classification adopted
by Agassiz wonderfully agree with the results of the less simple
principles adopted by Cuvier and the other masters in this department
of Natural History. Now, it is peculiar to yet a fourth order, the
Ganoidean, or shining-scaled order, that by much the greater number
of the genera which it comprises exist only in the fossil state. At
least five sixths of the whole were ascertained to be extinct several
years ago, at a time when the knowledge of fossil Ichthyology was
much more limited than at present: the proportions are now found to
be immensely greater on the side of the dead. And this order seems to
have included all the semi-osseous, semi-cartilaginous ichthyolites
of the Lower Old Red Sandstone: the enamelled scale is the
characteristic, according to Agassiz's principle of classification,
of the existences that filled the gap so often alluded to as existing
in the present creation. All their scales glitter with enamel: they
bore to this order the relation that the cartilaginous fish bear
to the Placoidean order, the thorny-finned fish to the Ctenoidean
order, and the soft-finned fish to the Cycloidean order. It also
included, with the semi-cartilaginous, the sauroid fish--those master
existences and tyrants of the earlier vertebrata; and both classes
find their representatives among the comparatively few ganoid fishes
of the present creation; the one in the sturgeon family, which of
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