The recent bony fishes are divided, according to the Cuvierian system
of classification, into two great orders, the soft-finned and the
thorny-finned order--the _Malacopterygii_ and the _Acanthopterygii_.
In the former the rays of the fins are thin, flexible, articulated,
branched: each ray somewhat resembles a jointed bamboo; with this
difference, however, that what seems a single ray at bottom,
branches out into three or four rays a-top. In the latter, (the
thorny-finned order,)--especially in their anterior dorsal, and
perhaps anal fins,--the rays are stiff continuous spikes of bone,
and each stands detached as a spear, without joint or branch. The
perch may be instanced as a familiar illustration of this order--the
gold-fish of the other. Now, between the fins of two sets--shall I
venture to say orders?--of the ichthyolites of the Lower Old Red
Sandstone, an equally striking difference obtains. The fin of the
_Osteolepis_, with its surface of enamelled and minutely jointed
bones, I have already described as a sort of bird-wing fin. The
naked rays, with their flattened surfaces, lay thick together as
feathers in the wing of a bird--so thick as to conceal the connecting
membrane; and fins of similar construction characterized the families
of the _Dipterus_, _Diplopterus_, _Glyptolepis_, _Cheirolepis_,
_Holoptychius_, and, I doubt not, many other families of the same
period, which await the researches of future discoverers. But the
fins of another set of ichthyolites, their contemporaries, may be
described as bat-wing fins: they presented to the water a broad
expanse of membrane; and the solitary ray which survives in each was
not a jointed, but a continuous spear-like ray. The fins of this set,
or order, are thorny-fins, like those of the _Acanthopterygii_; the
anterior edge of each, with the exception of, perhaps, the caudal
fin, which differs in construction from the others, is composed of a
strong, bony spike. Such, with some tacit reference, perhaps, to the
similar Cuvierian principle of classification, were the distinctive
differences, on the strength of which I used to arrange two of my
groups of fossils on the granitic boulder; and the influence of the
same principle, almost instinctively exerted,--for, in writing the
previous pages, I scarce thought of its existence,--has, I find,
given to each group its own chapter.
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