Instead of making their appearance singly and successively, and passing
imperceptibly one into another, all three groups of Vascular Cryptogams,
and the Gymnosperms into the bargain, come on the stage together, in the
Devonian strata; and Monocotyledons in the lower Carboniferous
immediately following. There is no trace whatever of the development of
any of these forms from the earlier cellular cryptogams:
But [says Mr. Carruthers] the evolution of the Vascular Cryptogams,
and the Phanerogams, from the green seaweeds, through the
liverworts and mosses, if it took place, must have been carried on
through a long succession of ages, and by an innumerable series of
advancing steps; and yet we find not a single trace either of the
early water forms or of the later and still more numerous dry-land
forms. The conditions that permitted the preservation of the
fucoids in the Llandovery rocks at Malvern, and of similar cellular
organisms elsewhere, were, at least, fitted to preserve _some_
record of the necessarily rich floras, if they existed, which
through immense ages, led by minute steps to the Conifer
[_Gymnosperm_] and Monocotyledon of these Palæozoic Rocks.
Further, these earliest plants are not generalized forms of the
various tribes to which they belong, but they are as highly
specialized as any subsequent representatives of the particular
group to which they belong, and wherever they differ from later
plants, it is in the possession of a more perfect organization.
* * * * *
From all which facts Mr. Carruthers thus argues:
The complete absence of intermediate forms, and the sudden and
contemporaneous appearance of highly organized and widely separated
groups, deprive the hypothesis of genetic evolution of any
countenance from the plant-record of these ancient rocks. The whole
evidence is against evolution, and there is none for it.[252]
Dicotyledons furnish evidence of especial value. On account of their
higher organization, they are easily distinguished from both
Monocotyledons and Gymnosperms; and they present features which clearly
differentiate them amongst themselves. They did not make their entry
till after a long interval--and their remains are therefore to be found
in strata comparatively recent and better known to us than those of the
older rocks. It is in the Chalk, the newest of the Secondary or Mesozoic
formations, that they first exhibit themselves, and they do it in the
same fashion as their predecessors.
When the Dicotyledons appear in the upper cretaceous beds,
representatives of the three great groups [_Apetalæ_, _Monopetalæ_,
_Polypetalæ_] appear together in the same deposit. Moreover, these
divisions are represented, not by generalized types, but by
differentiated forms, which, during the intervening epochs, have not
developed even into higher generic groups.
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