A singularly instructive field for the study of the mutability or
stability of species should be afforded by the lower forms of life, in
which organization is reduced to a minimum, they being mere masses of
protoplasm without even a containing envelope, while their nourishment
is of the simplest. It would therefore appear that environment should be
all-potent to modify them and produce specific modifications, while the
extreme rapidity with which they propagate their kind, and that
unisexually, ought to require no vast extent of time to make such
transmutations apparent.
It is found, however, on the contrary, that nowhere in organic nature
does the type remain more rigidly persistent. Professor Macbride, for
example, tells us,[327]
"The Myxomycetæ may be regarded as the organic group in which the forces
of heredity,--whatever these forces may be--are at their maximum: they
have responded as little as possible to the influence of their
environment."
To the same effect speaks Professor Paulesco of Bucharest, of other
elementary organisms.[328]
What is still more remarkable, these same organisms are extremely
sensitive to altered conditions of environment, which have a direct and
immediate influence, gravely modifying their morphological and
physiological characters, changes in respect of light, minute
alterations of temperature, or the introduction of a new chemical
substance, even in infinitesimal quantity, frequently causing them to
assume forms very different from the specific type, and profoundly
modifying their nutritive processes.
Here, it was at first thought, when Pasteur revealed their history, is
clear evidence of specific transformation. But he presently convinced
himself and others that it is not so, for although liable to assume such
polymorphic forms according to the conditions in which they find
themselves, there is no alteration of specific nature, and if the
original circumstances be restored, the original forms reappear--"une
élasticité functionelle de la cellule lui permettant de se plier à des
conditions variées d'existence sans changer d'être." (Pasteur.)
As M. Duclaux adds:[329]
"La notion d'espèce ne disparait pas pour cela. La variabilité est un
caractère comme un autre, bien que plus difficile à inscrire dans la
classification, et une espèce est aussi bien définie par les
sensibilités diverses qu'elle manifeste que par la petite liste des mots
et de propriétés dans laquelle on croyait pouvoir autrefois enfermer
toute son histoire.... La lien de l'espèce c'est la loi qui préside à
ces changements, et la variété des formes et des fonctions n'est pas du
tout en contradiction avec l'unité de l'espèce."
_B._ _Note on Chap. XV. p. 203._
Since the foregoing pages have been in type there has come to hand the
New York _Literary Digest_ of January 23, 1904, containing the following
article (p. 119).
"ARE THE DAYS OF DARWINISM NUMBERED?"
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