Recent investigations have led to the remarkable discovery that many
fern-like plants of the Carboniferous rocks, hitherto classed as
Cryptogams, were in reality seed-bearers, and thus intermediate between
Cryptogams and Cycads, the most primitive of existing seed-plants. They
have accordingly been placed in a special group "Cycadofilices," or
"Fern-Cycads," and regarded as transitional types, the view that they
are the remains of a natural bridge connecting the Ferns with the
Gymnosperms having received wide support,[330] and at first sight this
conclusion would appear natural and obvious. But here, as in other
cases, the difficulty is that the seeds which have been found are all
fully developed; there are none in the intermediate stages between true
spores and true seeds; we have the finished article, but no trace of
seeds in the making; which upon any theory of evolution must have been
exceedingly numerous. Hence Dr. Scott tells us:[331]
"The important discoveries of the seeds of the Pteridosperms scarcely
touch the question of descent, for these organs are of too advanced a
type to throw light on the probable derivation of the group."
In this instance, therefore, as in others, it remains true that in no
case is any trace found of rudimentary character in the earliest fossil
specimens of any class.
It is undoubtedly a further puzzle that some of the Carboniferous
cryptogams which did not bear real seeds, yet simulated them, a habit
not easily explained on evolutionary principles.
_D._ _The Course of Evolution._
The evidence of Professor Vines quoted in the text (pp. 202, 237)
receives a remarkable confirmation from that of Dr. Smith Woodward,
Keeper of Geology in the National Museum of Natural History. Speaking
before the International Congress of Arts and Science, St. Louis,
U.S.A., September 22nd, 1904, he thus touched upon the same question,
which he illustrated especially from the history of fossil fishes, which
he has made his special study.[332]
"It must be confessed that repeated discoveries have now left faint
hope that exact and gradual links will ever be forthcoming between
most of the families and genera. The 'imperfection of the record,'
of course, may still render some of the negative evidence
untrustworthy; but even approximate links would be much commoner in
collections than they actually are if the doctrine of gradual
evolution were correct. Palæontology, indeed, is clearly in favour
of the theory of discontinuous mutation, or advance by sudden
changes, which has lately received so much support from the
botanical experiments of H. de Vries.
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