The tendency of organic life, whether vegetable or animal, being to
propagate itself enormously,--and the life-sustaining capacity of the
earth being limited,--it necessarily follows that only a fraction of the
creatures which are born can survive to maturity, and that while those
best fitted to live will live, those less well fitted will die. Thus,
there is set up a constant struggle for existence, in which every
advantage, however slight, must tell, so that those possessing such
advantages in one generation will be the parents of the next. But in the
course of propagation, the offspring never exactly reproduce the parent
form, from which they vary, some in one way some in another, and as some
of these variations cannot help being advantageous to their possessors
in the struggle, we have here the required factor for the production of
new forms. Any thus beneficially equipped, (although the variation, and
consequently the advantage, must in each instance be exceedingly
slight,) will have the chances on their side against their less favoured
fellows, whom in the long run they will supplant. And as their
offspring, or some of them, will carry the profitable variation somewhat
further, the stream of life will thus be set in such a direction as will
ultimately bring about what might at first appear impossible
metamorphoses.
Thus, to take a simple and favourite illustration,[180] winged insects
inhabiting an island far from other land, are liable to be blown out to
sea and drowned. It is in consequence, an advantage to them to have
their power of flight curtailed, or taken away, and consequently in such
situations their wings are generally found to be so reduced as to permit
little or even nothing in the way of flying. Or to take an example of
another kind,[181] the extraordinary length of neck which characterizes
the giraffe enables it to browse on the higher branches of trees
inaccessible to other vegetable feeders, and thus gives it an advantage
over them in times of drought and scarcity of fodder. It can accordingly
be easily understood, how its present structure has resulted from
gradual elongations of the neck, each conferring on its possessor a
slight advantage.
Public-domain text, read in full here on John Shaqi.
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