The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th EditionDarwin, Charles
History
The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th Edition
Darwin, Charles
Evolution (Biology); Natural selection
It is difficult to imagine conditions of life more similar than deep
limestone caverns under a nearly similar climate; so that, in
accordance with the old view of the blind animals having been
separately created for the American and European caverns, very close
similarity in their organisation and affinities might have been
expected. This is certainly not the case if we look at the two whole
faunas; with respect to the insects alone, Schiödte has remarked: “We
are accordingly prevented from considering the entire phenomenon in any
other light than something purely local, and the similarity which is
exhibited in a few forms between the Mammoth Cave (in Kentucky) and the
caves in Carniola, otherwise than as a very plain expression of that
analogy which subsists generally between the fauna of Europe and of
North America.” On my view we must suppose that American animals,
having in most cases ordinary powers of vision, slowly migrated by
successive generations from the outer world into the deeper and deeper
recesses of the Kentucky caves, as did European animals into the caves
of Europe. We have some evidence of this gradation of habit; for, as
Schiödte remarks: “We accordingly look upon the subterranean faunas as
small ramifications which have penetrated into the earth from the
geographically limited faunas of the adjacent tracts, and which, as
they extended themselves into darkness, have been accommodated to
surrounding circumstances. Animals not far remote from ordinary forms,
prepare the transition from light to darkness. Next follow those that
are constructed for twilight; and, last of all, those destined for
total darkness, and whose formation is quite peculiar.” These remarks
of Schiödte’s it should be understood, apply not to the same, but to
distinct species. By the time that an animal had reached, after
numberless generations, the deepest recesses, disuse will on this view
have more or less perfectly obliterated its eyes, and natural selection
will often have effected other changes, such as an increase in the
length of the antennæ or palpi, as a compensation for blindness.
Notwithstanding such modifications, we might expect still to see in the
cave-animals of America, affinities to the other inhabitants of that
continent, and in those of Europe to the inhabitants of the European
continent. And this is the case with some of the American cave-animals,
as I hear from Professor Dana; and some of the European cave-insects
are very closely allied to those of the surrounding country. It would
be difficult to give any rational explanation of the affinities of the
blind cave-animals to the other inhabitants of the two continents on
the ordinary view of their independent creation. That several of the
inhabitants of the caves of the Old and New Worlds should be closely
related, we might expect from the well-known relationship of most of
their other productions. As a blind species of Bathyscia is found in
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