The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th EditionDarwin, Charles
History
The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th Edition
Darwin, Charles
Evolution (Biology); Natural selection
attribute this similarity in the enlarged stems of these three plants,
not to the vera causa of community of descent, and a consequent
tendency to vary in a like manner, but to three separate yet closely
related acts of creation. Many similar cases of analogous variation
have been observed by Naudin in the great gourd family, and by various
authors in our cereals. Similar cases occurring with insects under
natural conditions have lately been discussed with much ability by Mr.
Walsh, who has grouped them under his law of equable variability.
With pigeons, however, we have another case, namely, the occasional
appearance in all the breeds, of slaty-blue birds with two black bars
on the wings, white loins, a bar at the end of the tail, with the outer
feathers externally edged near their bases with white. As all these
marks are characteristic of the parent rock-pigeon, I presume that no
one will doubt that this is a case of reversion, and not of a new yet
analogous variation appearing in the several breeds. We may, I think,
confidently come to this conclusion, because, as we have seen, these
coloured marks are eminently liable to appear in the crossed offspring
of two distinct and differently coloured breeds; and in this case there
is nothing in the external conditions of life to cause the reappearance
of the slaty-blue, with the several marks, beyond the influence of the
mere act of crossing on the laws of inheritance.
No doubt it is a very surprising fact that characters should reappear
after having been lost for many, probably for hundreds of generations.
But when a breed has been crossed only once by some other breed, the
offspring occasionally show for many generations a tendency to revert
in character to the foreign breed—some say, for a dozen or even a score
of generations. After twelve generations, the proportion of blood, to
use a common expression, from one ancestor, is only 1 in 2048; and yet,
as we see, it is generally believed that a tendency to reversion is
retained by this remnant of foreign blood. In a breed which has not
been crossed, but in which _both_ parents have lost some character
which their progenitor possessed, the tendency, whether strong or weak,
to reproduce the lost character might, as was formerly remarked, for
all that we can see to the contrary, be transmitted for almost any
number of generations. When a character which has been lost in a breed,
reappears after a great number of generations, the most probable
hypothesis is, not that one individual suddenly takes after an ancestor
removed by some hundred generations, but that in each successive
generation the character in question has been lying latent, and at
last, under unknown favourable conditions, is developed. With the
barb-pigeon, for instance, which very rarely produces a blue bird, it
is probable that there is a latent tendency in each generation to
produce blue plumage. The abstract improbability of such a tendency
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