The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th EditionDarwin, Charles
History
The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th Edition
Darwin, Charles
Evolution (Biology); Natural selection
Mr. Mivart has taken up this case, and remarks that a sudden
spontaneous transformation in the position of the eyes is hardly
conceivable, in which I quite agree with him. He then adds: “If the
transit was gradual, then how such transit of one eye a minute fraction
of the journey towards the other side of the head could benefit the
individual is, indeed, far from clear. It seems, even, that such an
incipient transformation must rather have been injurious.” But he might
have found an answer to this objection in the excellent observations
published in 1867 by Malm. The Pleuronectidæ, while very young and
still symmetrical, with their eyes standing on opposite sides of the
head, cannot long retain a vertical position, owing to the excessive
depth of their bodies, the small size of their lateral fins, and to
their being destitute of a swimbladder. Hence, soon growing tired,
they fall to the bottom on one side. While thus at rest they often
twist, as Malm observed, the lower eye upward, to see above them; and
they do this so vigorously that the eye is pressed hard against the
upper part of the orbit. The forehead between the eyes consequently
becomes, as could be plainly seen, temporarily contracted in breadth.
On one occasion Malm saw a young fish raise and depress the lower eye
through an angular distance of about seventy degrees.
We should remember that the skull at this early age is cartilaginous
and flexible, so that it readily yields to muscular action. It is also
known with the higher animals, even after early youth, that the skull
yields and is altered in shape, if the skin or muscles be permanently
contracted through disease or some accident. With long-eared rabbits,
if one ear flops forward and downward, its weight drags forward all the
bones of the skull on the same side, of which I have given a figure.
Malm states that the newly-hatched young of perches, salmon, and
several other symmetrical fishes, have the habit of occasionally
resting on one side at the bottom; and he has observed that they often
then strain their lower eyes so as to look upward; and their skulls are
thus rendered rather crooked. These fishes, however, are soon able to
hold themselves in a vertical position, and no permanent effect is thus
produced. With the Pleuronectidæ, on the other hand, the older they
grow the more habitually they rest on one side, owing to the increasing
flatness of their bodies, and a permanent effect is thus produced on
the form of the head, and on the position of the eyes. Judging from
analogy, the tendency to distortion would no doubt be increased through
the principle of inheritance. Schiödte believes, in opposition to some
other naturalists, that the Pleuronectidæ are not quite symmetrical
even in the embryo; and if this be so, we could understand how it is
that certain species, while young, habitually fall over and rest on the
left side, and other species on the right side. Malm adds, in
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account