The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th EditionDarwin, Charles
History
The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th Edition
Darwin, Charles
Evolution (Biology); Natural selection
Mr. Mivart adduces this case, chiefly on account of the supposed
difficulty of organs, namely the avicularia of the Polyzoa and the
pedicellariæ of the Echinodermata, which he considers as “essentially
similar,” having been developed through natural selection in widely
distinct divisions of the animal kingdom. But, as far as structure is
concerned, I can see no similarity between tridactyle pedicellariæ and
avicularia. The latter resembles somewhat more closely the chelæ or
pincers of Crustaceans; and Mr. Mivart might have adduced with equal
appropriateness this resemblance as a special difficulty, or even their
resemblance to the head and beak of a bird. The avicularia are believed
by Mr. Busk, Dr. Smitt and Dr. Nitsche—naturalists who have carefully
studied this group—to be homologous with the zooids and their cells
which compose the zoophyte, the movable lip or lid of the cell
corresponding with the lower and movable mandible of the avicularium.
Mr. Busk, however, does not know of any gradations now existing between
a zooid and an avicularium. It is therefore impossible to conjecture by
what serviceable gradations the one could have been converted into the
other, but it by no means follows from this that such gradations have
not existed.
As the chelæ of Crustaceans resemble in some degree the avicularia of
Polyzoa, both serving as pincers, it may be worth while to show that
with the former a long series of serviceable gradations still exists.
In the first and simplest stage, the terminal segment of a limb shuts
down either on the square summit of the broad penultimate segment, or
against one whole side, and is thus enabled to catch hold of an object,
but the limb still serves as an organ of locomotion. We next find one
corner of the broad penultimate segment slightly prominent, sometimes
furnished with irregular teeth, and against these the terminal segment
shuts down. By an increase in the size of this projection, with its
shape, as well as that of the terminal segment, slightly modified and
improved, the pincers are rendered more and more perfect, until we have
at last an instrument as efficient as the chelæ of a lobster. And all
these gradations can be actually traced.
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