The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th EditionDarwin, Charles
History
The Origin of Species by Means of Natural Selection: Or, the Preservation of Favoured Races in the Struggle for Life, 6th Edition
Darwin, Charles
Evolution (Biology); Natural selection
and under different circumstances.
In the vegetable kingdom Mr. Mivart only alludes to two cases, namely
the structure of the flowers of orchids, and the movements of climbing
plants. With respect to the former, he says: “The explanation of their
_origin_ is deemed thoroughly unsatisfactory—utterly insufficient to
explain the incipient, infinitesimal beginnings of structures which are
of utility only when they are considerably developed.” As I have fully
treated this subject in another work, I will here give only a few
details on one alone of the most striking peculiarities of the flowers
of orchids, namely, their pollinia. A pollinium, when highly developed,
consists of a mass of pollen-grains, affixed to an elastic foot-stalk
or caudicle, and this to a little mass of extremely viscid matter. The
pollinia are by this means transported by insects from one flower to
the stigma of another. In some orchids there is no caudicle to the
pollen-masses, and the grains are merely tied together by fine threads;
but as these are not confined to orchids, they need not here be
considered; yet I may mention that at the base of the orchidaceous
series, in Cypripedium, we can see how the threads were probably first
developed. In other orchids the threads cohere at one end of the
pollen-masses; and this forms the first or nascent trace of a caudicle.
That this is the origin of the caudicle, even when of considerable
length and highly developed, we have good evidence in the aborted
pollen-grains which can sometimes be detected embedded within the
central and solid parts.
Public-domain text, read in full here on John Shaqi.
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