Quite recently Gerhard Holm has worked over again the very numerous
specimens of _Eurypterus Fischeri_, which are obtainable at Rootziküll, and
has thrown new light on the relation of the metastoma to the mouth-parts.
His preparations show clearly that the true lower lip of Eurypterus was not
the metastoma, for when the metastoma is removed another plate (_End._,
Fig. 105, B) situated {241}internally to it is disclosed, which, in his
view, corresponds to the sternite between the bases of the pro-somatic
appendages in Limulus, _i.e._ to the sternite called by Lankester, the
pro-mesosternite (_End._, Fig. 103). This inner plate formed with the
metastoma ((7) Fig. 105) and the ectognaths (6) a chamber closed
posteriorly, within which the bases of the ectognaths worked. In other
words, the removal of the metastoma discloses in Eurypterus the true
anterior ventral surface of the animal which corresponds to that of
Limulus, or of the scorpion group, with its pro-mesosternite and laterally
attached gnathites or sterno-coxal processes. To this inner plate or
pro-mesosternite Holm gives the name of _endostoma_.
To the anterior edge of the endostoma a thinner membrane is attached which
passes inwards in the direction of the throat, and forms, therefore, the
lower lip (_Hyp._, Fig. 105, B) of the passage of the mouth (_olf. p._).
This membrane bears upon its surface a tuft of hairs, which he thought were
probably olfactory in function. Consequently, in his preliminary
communication, he describes this lower lip as forming, in all probability,
an olfactory organ; in his full communication he repudiates this
suggestion, because he thinks it unlikely that such an organ would be
situated within the mouth. I feel sure that if Holm had referred to
Croneberg's paper, and seen how the true mouth in all the scorpion group is
situated at the base of an olfactory passage, he would have recognized that
his first suggestion is in striking accordance with the nature of the
entrance to the mouth in other scorpions.
Public-domain text, read in full here on John Shaqi.
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