On all the branchial appendages in Limulus, special sense-organs are found
of a most conspicuous character. They form in the living animal bluish
convex circular patches, the situation of which on the appendages is shown
in Fig. 58. These organs are not found on the non-branchial operculum.
Gegenbaur, who was the first to describe them, has pointed out how the
surface of the organ is closely set with chitinous goblets shaped as seen
in Fig. 144, A, which do not necessarily project free on the surface, but
are extruded on the slightest pressure. Each goblet fits into a socket in
the chitinous covering, and is apparently easily protruded by variations of
pressure from within. The whole surface of the organ on the appendage is
slightly bulged in the living condition, and the chitin is markedly softer
here than in the surrounding part of the limb. Each of these organs is
surrounded by a thick protection of strongly branching spines. On the
surface of the organ itself no spines are found, only these goblets, so
that the surface-view presents an appearance as in Fig. 144, B. Each goblet
possesses a central pore, which is the termination of a very fine, very
tortuous, very brittle chitinous tubule (_ch.t._), which passes from the
goblet through the layers of the chitin into the subjacent tissue. The
goblets vary considerably in size, a few very large ones being scattered
here and there. The fine chitinous tubule is especially conspicuous in
connection with these largest goblets. In the smaller ones there is the
same appearance of a pore and a commencing tube, but I have not been able
to trace the tube through the chitinous layers, as in the case of the
larger goblets.
{360}[Illustration: FIG. 144.--A, A GOBLET FROM ONE OF THE BRANCHIAL
SENSE-ORGANS OF LIMULUS (_ch.t._, chitinous tubule); B, SURFACE VIEW OF A
PORTION OF A BRANCHIAL SENSE-ORGAN.]
[Illustration: FIG. 145.--THE ENDOGNATHS OF LIMULUS PUSHED OUT OF THE WAY
ON ONE SIDE IN ORDER TO SHOW THE POSITION OF THE FLABELLUM (_fl._)
PROJECTING TOWARDS THE CRACK BETWEEN THE PROSOMATIC AND MESOSOMATIC
CARAPACES.]
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