_bl._, blood-spaces; _n._, nerve; _gl._, layer of ganglion-cells; _p._,
pigment layer; _ch._, 1, 2, 3, the three layers of chitin; _ch.t._,
chitinous tubule in large tube of sense-organ; _cap._, capitellum or
swollen extremity of large tube; _can._, very fine porous canals or
canaliculi of chitin.]
In Fig. 148 I give a magnified representation of a section through three of
these flabellar sense-organs. As is seen, the section divides itself into
four zones: (1) the chitinous layer (_ch._); (2) the layer of pigment
(_p._) and hypodermal cells; (3) the layer of ganglion-cells (_gl._); and
(4) the layer of nerve-fibres (_n._) and blood-spaces (_bl._). The
chitinous layer is composed of the usual three zones of the Limulus
surface--externally (Fig. 148), a thin homogeneous layer, followed by a
thick layer of chitin (3), in which the fine vertical tubules or canaliculi
are well marked; the external portion (2) of this layer is differentiated
from the rest by the presence of well-marked horizontal layers in addition
to the canaliculi.
In this chitinous layer the special sense-organs are found. They consist of
a large tube which passes through all the layers of the chitin except the
thin homogeneous most external layer. {364}This tube is conical in shape,
its base, which rests on the pigmented layer, being so large and the organs
so crowded together that a section of the chitin across the base of the
tubes gives the appearance of a honeycomb, the septa of which is all that
remains of the chitin. This large tube narrows down to a thin elongated
neck as it passes through the chitin, and then, at its termination, bulges
out again into an oval swelling (_cap._) situated always beneath the
homogeneous most external layer of chitin. Within this tube a fine
chitinous tubule (_ch. t._) is situated similar to that seen in the
branchial sense-organs; it lies apparently free in the tube, not straight,
but sinuous, and it passes right through all the chitinous layers to open
at the surface as a pore; in the last part of its course, where it passes
through the most external layer (1) of chitin, it lies always at right
angles to the surface.
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