The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
Forty-three specimens are mounted to exemplify the variations in size
and characters of cervical vertebræ. These for the most part are
specific characters; and between the axis and the first dorsal vertebra
the variations in an individual were slight. [Those nearest to the
back, as in birds, are widest in front, and have the highest neural
arches.] The associated series show commonly four cervical vertebræ
behind the axis, and in two cases apparently five; never more. So that
as seven appears to be the number of true cervical vertebræ in most
if not all of the German Pterodactyles, it may be presumed that the
Cretaceous Ornithosaurians also had this character in common, with
Mammals, and probably as persistent. In Iguana there are 6, in Monitor
7, and in Crocodile 8.
The centrum is united to the neural arch as in birds, without a trace
of suture; sometimes the neural arch is no wider than the centrum,
sometimes it extends over the centrum on each side. Those forms
with a narrow neural arch have the neural spine high, and its sides
look forward as well as outward. The pneumatic foramen is oblique.
An example is figured by Prof. Owen, in the memoir on Pterodactylus
simus, pl. 2, fig. 4. The forms with a wide neural arch have the neural
spine rising from the middle of the dorsal surface, erect and equally
compressed from side to side. The pneumatic foramen is horizontal. An
example is figured in Prof. Owen's memoir on Pt. simus, pl. 2, fig. 1.
These two forms of cervical vertebræ may be regarded as typifying two
genera.
In both forms many characters occur in common, and as the specimens
illustrative of special modifications will be described hereafter,
the following description has been made to embrace the chief
characteristics of these vertebræ in Cretaceous Ornithosaurians.
The inferior aspect of the centrum is oblong (being narrower than
long), or quadrate; when quadrate the additional lateral expansions are
external to the pneumatic foramina, and are formed by the neural arch
and zygapophyses. The centrum proper is a little wider in front than
behind, and the side outlines are concave. The base of the centrum is
flattened, or more or less hollow, or more or less tumid and regularly
convex; in front there is often a mesial ridge, which never reaches
the posterior articulation, and forms a prominent tubercle at the base
of the anterior articulation. At the posterior end the outline of the
centrum is concave, and mesially the bone has a hollow corresponding
to the tubercle in front of the adjacent vertebra; and the part of
the centrum on each side is prolonged slightly into a strong rounded
or flattened tubercle below the side borders of the posterior
articulation; these posterior processes, in vertebræ in situ, fitted,
on each side of the mesial anterior process of the vertebra behind, on
to concavities more or less marked. Analagous processes are developed
in the cervical vertebræ of many birds.
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