The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
_The base of the skull._ The bones here indicated are the
basi-occipital, basi-temporal, and basi-sphenoid. The former two
have come away as from an articular joint, and are wanting. The
basi-occipital does not enter into the floor of the cranial cavity, and
only rims the foramen magnum. But its basi-temporal expansion rests
beneath the posterior part of the basi-sphenoid forming the base of the
skull; its long convex anterior end fits into the concave groove at the
back of the anterior part of the sphenoid. The squamous basi-temporal
bone appears in this species to have been as long as the foramen magnum
is wide, and to have been relatively thicker than in the other form
already described.
The _basi-sphenoid_ is a thin expanded bone forming the floor for the
cerebellum, and terminating anteriorly in a triangular mass, while the
slightly convex part behind, covered with the basi-temporals, is nearly
square. It enters into the foramen magnum, forming its lower part;
and is confluent with the ex-occipitals behind, with the periotic,
alisphenoid and perhaps with the squamosal at the side; and as in birds
all these sutures are obliterated. This is probably the only instance
in the Animal Kingdom in which the basi-sphenoid takes so important
and singular a share in the functions of the basi-occipital bone. The
anterior part of the basi-sphenoid projects below the posterior part,
is nearly flat on the basal surface, and forms an equilateral triangle
with the apex in front and base behind. In the middle of the triangular
bone is a slight longitudinal ridge, and behind the middle of each
outer side a rather large foramen which appears to be the inferior
opening for the carotid artery. The triangular part is hollow and as
long as the quadrate portion. The lateral parts of this anterior bone
are nearly flat. They converge upwards and are rounded in front to
form the boundary of the pituitary fossa, and do not appear to have
terminated in a spine. Above are the alisphenoids.
_The upper part of the skull_ is divided into two segments by a strong
straight transverse ridge, which leaves the occipital bones behind, and
the parietal &c. in front.
The occipital bones anchylosed together are about two-thirds the width
of the foramen magnum, and of the parietal bones, with which latter the
supra-occipital makes an angle of 45°. The surface is irregular, and
especially is marked by a deep concavity just above each ex-occipital.
The supra-occipital projects slightly over the plane of the foramen
magnum, to which the strong ridge bounding the segment in front is
parallel. The great foramen is nearly round, being slightly compressed
at the upper part of the sides: it measures 3/8 of an inch high and is
nearly as wide.
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