The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
The only portion of the specimen now to be described is the large
region at each side looking downward, which extends from the occipital
ridge to the sphenoid. It is an irregular pentangular hollow with many
cavities, the hinder of which are for the ear. Two cavities above
these, under the widest part of the skull, appear to be a double
articulation for the quadrate bone. The outer transverse one with
the squamosal is separated by a deep groove from the inner and more
vertical one, which may therefore be regarded as with the petrosal
bone. These excavations form the posterior half of the pentagon. The
anterior half is a smooth rhombus not separable from the basi-sphenoid.
Such is the external appearance of the occipital and parietal segments
of the skull of a Cambridge Pterodactyle. Each segment forms a large
ring of thin bone, inclosing part of a brain-cavity as large as that
of a bird and shaped like that of a bird; and which moreover is made
up of the same bones as the cranium of a bird; and these are in almost
exactly the same proportions as those of the Common Cock.
My own investigations do not substantiate Wagner's discovery, that the
back part of the skull resembles that of the Monitor. Iguana would
have offered a slightly nearer comparison, but they both differ from
Cambridge specimens of Pterodactyles in characters like these.
In the lizard,
The cranial bones do not enclose the brain.
There is no division of the back of the skull into an occipital
segment and a parietal segment by a girdling crest.
The squamosal bone does not enter into the cranial wall.
The quadrate bone does not articulate with the wall of the brain-case.
While the peculiar backward development of wings of the parietal in a
diverging V form, give the Lizard skull an aspect of its own.
So that it must be asserted that the differences of these Pterodactyles
from Lizards are so wide as to preclude comparison.
With the Crocodile, in which the cranial bones are massive, and the
quadrate bone firmly packed in the skull, comparison would be no less
difficult.
The Delphinidæ, in both the form of the jaws and of the back of the
head, give some support to Wagler's fancy, in putting the Pterodactyle
into his curious creation, the Gryphi[T]. But in the porpoises the
parietal bones form as narrow a band as they do in the Duck; and are
quite unlike the bones here described. In the Dolphin the two condyles
almost unite into one semicircular condyle (in young specimens), owing
to the enormous development of the ex-occipitals, which almost if
not entirely exclude the basi-occipital from the foramen magnum. The
dolphin moreover has no quadrate bone. But notwithstanding the absence
of a division into occipital and parietal segments, the form and
arrangement of the bones in the skull of the porpoises approximate more
to the Cambridge Pterodactyles than is the case with Lizards.
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