The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of CambridgeSeeley, H. G. (Harry Govier)
Science
The Ornithosauria: An elementary study of the bones of Pterodactyles made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge
Seeley, H. G. (Harry Govier)
Pterosauria
Which, if the fore-limbs were kept together as in ordinary quadrupeds,
would give a height to the body of about 2 ft. 6 in., but as the limbs
probably spread in walking as among the bats, the hind-limb would give
a better idea of the height of the animal.
(Flesh, sacrum, os innominatum). 2 in.
Femur 4 "
(_Tibia_) 6 "
(Metatarsus, &c.) 1 "
Which would give a height of about 13 inches; and, standing in the
position of a bird, the height to the crown of the head would be about
2 feet. The majority of the Ornithosaurians of the Cambridge Greensand
are of this size.
The spread of the wings, if there were 4 phalanges, would be
Body 10 in. wide.
Two arms 5 ft. 2 "
Two wing fingers 7 0 "
Giving a total expanse of about 13 feet. But, from the indications
of the wing-finger, I should incline to think an expanse of 10 feet
a truer estimate. The largest species attained to twice this size,
and the smallest was a fourth as large. Another memoir will present
descriptions and restorations of the Greensand species.
Habits.
The varying organization of different Ornithosaurians probably depends
on the different habits of the tribes. That they could all fly is
probable from the enormous radial crest to the humerus and the great
development of the wing-bones, to which a wing-membrane was stretched,
comparable to that of a Bat in texture, but more comparable to a
Bird in its extent. The groups with long hind-legs probably had the
membrane limited to the bones of the arm, while in the species with
small hind-legs it may have attained even as great a development as in
Bats, though there is no reason for suspecting that it extended to
the tail. A Pterodactyle cannot be supposed to have hung itself up by
the hind-legs as does a Bat, because the hind-claws appear invariably
to be directed forward. A Bat walks upon four legs with considerable
elegance and speed; the wing is folded in, close to the side, so as to
be scarcely noticed; and the outer claw is free to climb with. There
can be little doubt but that Pterodactyles walked in a similar way. The
thickened mammilate knob at the proximal end of the first phalange is
well calculated for contact with the ground. And if it were supposed
that the large wing-metacarpal bone were only used to support the wing,
and the small metacarpals only used to support the claws by which
the creature has sometimes been pictured suspending itself, it would
be difficult to believe that the forces of pressure and tension in
flying so exactly corresponded to the forces manifested in suspension
as to cause the large and the small metacarpals invariably to attain
the same length. A correspondence of this kind may be presumed to
indicate a correspondence in function; and since the animal did not
fly by means of its claws, the inference is that it walked by means of
the metacarpal bones.
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