The arboreal life had many other consequences. It led to an increased freedom of movement of the thigh on the hip joint, to muscular arrangements for balancing the body on the leg, to making the backbone a supple yet stable curved pillar, to a strongly developed collar-bone which is only found well-formed when the fore-limb is used for more than support, and to a power of "opposing" the thumb and the big toe to the other digits of the hand and foot--an obvious advantage for branch-gripping. But the evolution of a free hand made it possible to dispense with protrusive lips and gripping teeth. Thus began the recession of the snout region, the associated enlargement of the brain-box, and the bringing of the eyes to the front. The overcrowding of the teeth that followed the shortening of the snout was one of the taxes on progress of which modern man is often reminded in his dental troubles. Another acquisition associated with arboreal life was a greatly increased power of turning the head from side to side--a mobility very important in locating sounds and in exploring with the eyes. Furthermore, there came about a flattening of the chest and of the back, and the movements of the midriff (or diaphragm) came to count for more in respiration than the movements of the ribs. The sense of touch came to be of more importance and the sense of smell of less; the part of the brain receiving tidings from hand and eye and ear came to predominate over the part for receiving olfactory messages. Finally, the need for carrying the infant about among the branches must surely have implied an intensification of family relations, and favoured the evolution of gentleness. [Illustration: _Photo: New York Zoological Park._ THE GIBBON IS LOWER THAN THE OTHER APES AS REGARDS ITS SKULL AND DENTITION, BUT IT IS HIGHLY SPECIALIZED IN THE ADAPTATION OF ITS LIMBS TO ARBOREAL LIFE] [Illustration: _Photo: New York Zoological Park._ THE ORANG HAS A HIGH ROUNDED SKULL AND A LONG FACE] [Illustration: _Photo: British Museum (Natural History)._ COMPARISONS OF THE SKELETONS OF HORSE AND MAN Bone for bone, the two skeletons are like one another, though man is a biped and the horse a quadruped. The backbone in man is mainly vertical; the backbone in the horse is horizontal except in the neck and the tail. Man's skull is mainly in a line with the backbone; the horse's at an angle to it. Both man and horse have seven neck vertebrae. Man has five digits on each limb; the horse has only one digit well developed on each limb.]