But acquired characters were not really transmitted after all. So
those who clung to Weismannism argued--an unnecessary task surely
if this transmissibility were inconceivable. We cannot discuss the
evidence here, and it is unnecessary that we should do so, since it is
all considered in the popular books on heredity. There is an apparent
consensus of opinion in these books which should not influence the
reader unfamiliar with zoological literature, nor obscure the fact
that many zoologists and botanists accept the opposite conclusion.
The discussion is all very tiresome, but we may glean some results of
positive value from it. It is unquestionable that very few conclusive
and adequate investigations have been made: one cannot help noticing
that the literature contains an amount of controversy out of all
proportion to the amount of sound experimental and observational
work actually carried out. Most of the experiments deal with the
consideration of traumatic lesions or mutilations, and it seems to be
proved that such defects are not transmitted, or at least are very
rarely transmitted. The tails of kittens have been cut off; the ears
of terrier-dogs have been lopped; and the feet and waists of Chinese
and European ladies have been compressed, and all throughout very
numerous generations, yet these defects are not transmitted from parent
to offspring. This kind of evidence forms the bulk of that which
orthodox zoological opinion has adduced in favour of the belief in
the non-inheritability of acquired characters, but does it all really
matter? What might be transmitted is a useful, purposeful modification
of morphology, or functioning, or behaviour, induced by the environment
throughout a number of generations--an adaptation rather than a harmful
lesion. There is little conclusive evidence that such adaptations
are inherited, though anyone who carefully studies the evidence in
existence will not be likely to say that they are certainly not
transmitted. Does, for instance, the blacksmith transmit his muscular
shoulders and arms to his sons, or the pianiste her supple wrists and
fingers to her daughters? There are no observations and experiments in
the literature worthy of the importance attaching to the question at
issue.
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