Surely the organism itself selects the variations of structure and
functioning that are exhibited by itself. If we hesitate to say
that these modifications are creations, let us say that they are
permutations of elements of structure, and that they were potential
in the organisation of the creature exhibiting them. They occur in
the latter if we must not say that they are produced. If they are
detrimental, the organism is the less able to live and reproduce, and
if it does reproduce, its progeny are subject to the same disability.
If, as is usual, they simply do not matter, they may or may not affect
the direction of evolution. If they are of advantage, that is, if they
confer increased mastery over the environment, over the inert things
with which the organism comes into contact, the latter enlarges its
universe or environment, lives longer, and transmits to its progeny its
increased powers of action. Indefinite increase of power over inert
matter is potential in living things, and variation converts this
potentiality into actuality.
This discussion is all very formal, but two conclusions emerge from it:
(1) the insufficiency of the mechanistic hypotheses of transformism
to account for all the diversity of life that has appeared on the
earth during the limited period of time which physics allows for the
evolutionary process. There does not appear to be any possibility of
meeting this objection if we continue to adhere to the hypothesis
of transformism already discussed: it faces us at every turn in our
discussion. How great a part is played, for instance, by “pure chance”
in the elimination of individual organisms during the struggle for
existence! Let us think of a shoal of sprats on which sea-birds are
feeding: it is chance which determines whether the birds prey on one
part of the shoal rather than another. Or let us think of the millions
of young fishes that are left stranded on the sea-shore by the receding
tide: it is chance that determines whether an individual fish will be
left stranded in a shallow sandpool which dries up under the sun’s
rays, rather than in a deeper one that retains its water until the tide
next flows over it. It is no use to urge that there is no such thing
as “pure chance,” and that what we so speak of is only the summation
of a multitude of small independent causes. Let us grant this, and
it still follows that the alternative of life or death to multitudes
of organisms depends not upon their adaptability but upon minute
un-co-ordinated causes which have nothing to do with their morphology
or behaviour. These are instances among many others which will occur
to the field naturalist: they shorten still further the time available
for natural selection in the shaping of species, for they reduce the
material on which this factor operates.
Public-domain text, read in full here on John Shaqi.
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