Thus there may be suppression of parts leading to entire disappearance
or to mere vestiges of the original morphology. A structure
degenerating through disuse may become removed from its typical
relations with other structures and may acquire altogether new ones. Or
its increasing importance may lead to its hypertrophy and to increased
complexity of structure, and perhaps to the inclusion of new anatomical
elements, or to the incorporation of other parts, the function of which
may originally have been quite different. In all sorts of ways organs
and organ-systems may become anatomically different as the result of
adaptive modifications, or indirectly as non-adaptive modifications
induced by the adaptive modifications of adjacent parts. It is the task
of comparative anatomy to trace these changes of morphology, aided
by the study of embryology and by the comparison of the structure of
the parts of fossil animals. Regarding the process of transformism as
proved by experiments and observations in breeding and heredity, the
naturalist endeavours to trace the lines along which evolution has
proceeded from the results of morphological investigations.
Such results cannot have more than a very limited value, and it is
often the case that several interpretations of morphological results
are equally probable. We may conclude that the existing Teleost and
Elasmobranch fishes are descended from a common stock which no longer
exists; we may similarly conclude that the Birds and Reptiles are
closely allied, more so than either group is to the Mammals; and we
may conclude that the Primates--the group of Mammals to which Man
belongs--is descended from some group allied to the existing Ungulates
or Insectivores, while the Mammals themselves may have come down
from some group of vertebrates related to both the Amphibia and the
Reptiles. But as to the nature of the animals which combined the
characters of the Birds and Reptiles, or of the Reptiles and Amphibia,
we know nothing. Palæontology, if its results were more numerous than
they are, would afford us the material for the discovery of these
“missing links,” and there can be no doubt that as the world becomes
better known our knowledge of palæontological stages in the history of
existing groups will become more complete, so that we may, in time,
possess an actual historical record of the phylogeny of the main groups
of animals. But it is remarkable that while the results of comparative
anatomy and embryology, aided by those of palæontology, enable us to
trace back short series of stages in the evolutionary process, they
still show us gaps at all the places where lines of descent ought to
converge. They show us, for instance, that the oldest Birds known were
decidedly reptilian in their morphology, but they do not show us an
animal which was neither Bird nor Reptile, but from which both groups
of Vertebrata have descended; and this is almost always the case in
our hypothetical schemes of phylogeny.
Public-domain text, read in full here on John Shaqi.
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