Are the arbitrary “stages” of the embryologists--the ovum, blastula,
gastrula, etc., phases in a system in the above sense, the only sense
in which the process can be regarded as capable of physico-chemical
analysis? What precisely is the embryo at the close of the process of
segmentation? It is an harmonious equipotential system, that is to say,
an assemblage of discrete organic parts or cells, each of which has all
the potentialities that every one of the others has. _Any_ cell in the
blastula may become a cell, or a series of such, in _any_ part of the
gastrula or pluteus larva. This is what the parts are in potentiality,
but actually their individual fates are different. The system is an
harmonious one, and each of its parts, although able to do whatever
any other part can do, yet does one thing only: it becomes an endoderm
cell, or an ectodermal cell, or a part of the skeleton, and so on; what
it does depends on its position with regard to the other cells. An
extensive manifoldness or diversity is produced, but this was not the
consequence of a preceding extensive manifoldness, for in the preceding
stage _all the parts of the system were the same_. The manifoldness of
the ovum or blastula--that potential manifoldness which became actual
in development--must be an intensive manifoldness, and admitting this
we must abandon the comparison of the ontogenetic (and, of course,
phylogenetic) processes with the phases of a physico-chemical system
in process of transformation. _Evolution is the transformation of an
intensive into an extensive manifoldness._
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