The typical activities of the chlorophyll-containing organism are of
this nature; they are anabolic. The organism may be a green land-plant;
a marine green, red, or brown alga; a yellow-green diatom, a yellow,
green, red, or brown peridinian or other holophytic protozoan; an
ascidian, mollusc, echinoderm, polyzoan, worm, or coral containing
“symbiotic algæ” (that is the chlorophyll-containing cells of some
plant organism which have become associated with the animal and
incorporated in its tissues). In all these cases the presence of
this chlorophyllian substance confers on the organism the power of
effecting the compensatory energy-transformation, by the aid of
which carbon dioxide and water are built up into starch. What this
transformation is, and what are the steps by which the carbon dioxide
and water become carbohydrate we do not exactly know. Solar radiation
impinging upon an inorganic substance is partly reflected and partly
absorbed. The absorbed fraction may become transformed in such a way
as to render the substance phosphorescent, or it may transform into
chemical energy, as when light impinges on a photographic plate, but
as a general rule it is transformed into heat. In the green plant,
however, the transformation of radiation into heat does not occur--at
least the heating is very small--and it passes directly or indirectly
into the potential chemical energy of the starch which is synthesised.
We must regard this power of absorbing radiation and utilising it in
compensatory transformations as a general character of protoplasm.
It is true that it is now specialised in the cells containing the
chlorophyll bodies, but there are indications that it may be present in
the tissues of the animal devoid of chlorophyll.
Other anabolic transformations occur in the animal. The food-stuffs
which are absorbed from the intestine are substances which have
undergone dissociations, the nature of which is such as to render them
capable of absorption and of reconstruction. These anabolic changes in
the higher animal are exceptional, and their usefulness lies in the
fact that by their means substances become capable of being transported
by the tissue fluids of the body.
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