The Physical Basis of Mind: Being the Second Series of Problems of Life and Mind.Lewes, George Henry
Philosophy
The Physical Basis of Mind: Being the Second Series of Problems of Life and Mind.
Lewes, George Henry
Mind and body; Psychophysiology
123. If apolar cells are embryonic forms of cells which afterwards
become multipolar, this interpretation will not suffice for the
unipolar cells. They are not only abundant, but are mature forms in
some organs, and in some animals; though in some organs they may truly
be regarded as embryonic. Thus in the human embryo up to the fourth
month all the cells of the spinal cord are said to be unipolar,[144]
later on they become multipolar. But in birds, rabbits, dogs, and
even man, the cells in the spinal ganglia are mainly (if not wholly)
unipolar;[145] nor is there any difficulty in observing the same fact
in the œsophageal ganglia of molluscs (see Fig. 22).
Such are the observations. They have indeed been forced into agreement
with the bipolar postulate, by the assumption that the single process
branches into two, one afferent, the other efferent.[146] But before
making observation thus pliant to suit hypothesis, it would be well to
look more closely into the evidence for the hypothesis itself. For my
own part, I fail to see the justification of the postulate; whereas
the existence of unipolar cells is an observation which has been amply
verified.
[Illustration: Fig. 18.--_Supposed union of two nerve-cells and a
fibre._ The processes subdivide into a minute network, in which the
fibre also loses itself.]
124. Bipolar cells abound; multipolar cells are still more abundant;
and these are the cells found in the gray substance of the neural
axis. Deiters, in his epoch-making work,[147] propounded an hypothetic
_schema_ which has been widely accepted. Finding that the large
cells in the anterior horn of the spinal cord gave off processes
of different kinds, one branched, the other unbranched, he held
that the latter process was the origin of the axis cylinder of
a nerve-fibre, whereas the branched process was protoplasm which
divided and subdivided, and formed the connection between one cell
and another. Gerlach has modified this by supposing that the minute
fibrils of the branching process reunite and form an axis cylinder
(Fig. 18). There is no doubt that some processes terminate in a fine
network; and there is a probability (not more) that the unbranched
process is always continuous with the axis cylinder of a motor nerve,
as we know it sometimes is with that of a dark-bordered fibre in the
white substances. This, though probable, is, however, very far from
having been demonstrated. Once or twice Kölliker, Max Schultze, and
Gerlach have followed this unbranched process as far as the _root_ of
a motor nerve; and they infer that although it could not be traced
further, yet it did really join an axis cylinder there. In support
Of this inference came the observations of Koschennikoff,[148] that
in the cerebrum and cerebellum, processes were twice seen continuous
with dark-bordered nerve-fibres. But the extreme rarity of such
observations amid thousands of cells is itself a ground for hesitation
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