The Physical Basis of Mind: Being the Second Series of Problems of Life and Mind.Lewes, George Henry
Philosophy
The Physical Basis of Mind: Being the Second Series of Problems of Life and Mind.
Lewes, George Henry
Mind and body; Psychophysiology
130. Besides cells and fibres, there is the _amorphous substance_,
which constitutes a great part of the central tissue, and also enters
largely into the peripheral tissue. It consists of finely granular
substance, and a network of excessively delicate fibrils, with nuclei
interspersed. Its character is at present _sub judice_. Some writers
hold it to be nervous, the majority hold it to be simply one of
the many forms of connective tissue: hence its name neuroglia, or
nerve-cement.
In the convolutions of the frozen brain Walther finds the cells and
fibres imbedded in a structureless semi-fluid substance wholly free
from granules; the granules only appear there when cells have been
crushed. It is to this substance he attributes the fluctuation of
the living brain under the touch, like that of a mature abscess; the
solidity which is felt after death is due to the coagulation of this
substance. Unhappily we have no means of determining whether the
network visible under other modes of investigation is present, although
invisible, in this substance. The neuroglia, as it appears in hardened
tissues, must therefore be described with this doubt in our minds.
If we examine a bit of central gray substance where the cells and
fibres are sparse, we see, under a low power, a network of fibrils
in the meshes of which lie nerve-cells. Under very high powers we
see outside these cells another network of excessively fine fibrils
embedded in a granular ground substance, having somewhat the aspect of
hoar-frost, according to Boll. It is supposed that the first network
is formed by the ultimate ramifications of the nerve-cell processes,
and that the second is formed by ramifications of the processes of
_connective_ cells. In this granular, gelatinous, fibrillar substance
nuclei appear, together with small multipolar cells not distinguishable
from nerve-cells except in being so much smaller. These nuclei are
more abundant in the tissue of young animals, and more abundant in the
cerebellum than in the cerebrum. The granular aspect predominates the
fresher the specimen, though there is always a network of fibrils; so
that some regard the granules as the result of a resolution of the
fibrils, others regard the fibrils as the linear crystallization (so to
speak) of the granules.[157]
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