The physiology of digestion considered with relation to the principles of dieteticsCombe, Andrew
Science
The physiology of digestion considered with relation to the principles of dietetics
Combe, Andrew
Diet; Digestion
In addition to the ordinary longitudinal and transverse fibres, the
colon presents three remarkable muscular bands running along its whole
length, and one of which is represented on the colon in the figure on
page 155. On the rectum all the three bands are seen. It is in the
colon and rectum that the feculent matter accumulates before it is
thrown out of the bowels, and these bands are useful chiefly by adding
to their propelling power.
The natural tendency of muscular fibre being to contract it may
naturally be supposed that, after the intestine is emptied, its
opposite sides will come into contact, and, by thus obliterating the
cavity altogether, present an obstacle to the subsequent passage
of any solid matter. But on inspecting the abdomen after death, we
rarely meet with any considerable portion thus contracted; and in
general, the whole intestines are distended to a greater or less
degree, according to circumstances. The agent by which this effect is
brought about, is one known more familiarly by the inconveniences and
pain to which it gives rise when in excess, than by its proper uses,
which are nevertheless important. I allude to the presence of air in
the bowels, which is as necessary to their healthy action as their
muscular contraction itself. Air, in fact, by its expansive energy,
forms the antagonist power to the muscular coat, and serves to dilate
the bowel after the requisite contraction has propelled its contents. A
certain degree of distention, indeed, not only is a stimulus to farther
muscular contraction, but is useful in facilitating the passage of the
subsequent portions of the feculent matter; and hence the injection
of air into the bowels in large quantity, has lately been employed
successfully in overcoming obstinate constipation.
The _mucous_, _internal_, or _villous_ coat of the intestine, also
resembles in many respects that of the stomach. It is a soft velvety
membrane, full of wrinkles or folds through the greater part of its
course, by means of which its surface is greatly increased in extent,
so as to afford ample space for the ramification of the bloodvessels,
nerves, and absorbents, with which it is very plentifully supplied. The
cut on p. 172 will convey some idea of its appearance, as seen in the
smaller intestine. So far as nutrition is concerned, the mucous coat is
the truly _essential_ part of the bowel. It alone is in direct contact
with the chyme, and in its cavity the bile and pancreatic juice perform
their respective parts, and give rise to the formation of chyle,
which is afterwards transmitted from its surface into the general
system. The peritoneal and muscular coats are useful only in affording
protection, and communicating the power of propelling its contents.
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