The Popular Science Monthly, August, 1900: Vol. 57, May, 1900 to October, 1900Various
Science
The Popular Science Monthly, August, 1900: Vol. 57, May, 1900 to October, 1900
Various
Science -- Periodicals; Technology -- Periodicals
Weismann[N] more recently accepts the view of Romanes, Morgan and
Lankester on the inadequacy of panmixia to explain the whole phenomena
of degeneration, and in his ‘Germinal Selection’ rejects the idea
of reversed selection, and suggests a new explanation for what
Romanes attributed to the failure of heredity and the Lamarckians to
transmission of the effects of disuse. The struggle of the parts of
Roux has been crowded by him back to the representatives of these parts
in the germ.
[N] The Monist, 1896, pp. 250-274.
“The phenomena observed in the stunting, or degeneration, of parts
rendered useless ... show distinctly that ordinary selection, which
operates by the removal of entire persons--personal selection, as I
prefer to call it--can not be the only cause of degeneration, for
in most cases of degeneration it can not be assumed that slight
individual vacillations in the size of the organ in question have
possessed selective value. On the contrary, we see such retrogressions
effected apparently in the shape of a continuous evolutionary process
determined by internal causes, in the case of which there can be no
question whatever of selection of persons or of a survival of the
fittest--that is, of individuals with the smallest rudiments. The
gradual diminution continuing for thousands and thousands of years
and culminating in its final and absolute effacement” can only be
accomplished by germinal selection. Germinal selection as applied to
degeneration is the formal explanation of Romanes’ failure of heredity
through the struggle of parts for food. “Powerful determinants will
absorb nutriment more rapidly than weaker determinants. The latter,
accordingly, will grow more slowly and will produce weaker determinants
than the former.” If an organ is rendered useless, the size of this
organ is no longer an element in personal selection. This alone would
result in a slight degeneration. Minus variations are, however,
supposed to rest “on the weaker determinants of the germ, such as
absorb nutriment less powerfully than the rest. This will enable the
stronger determinants to deprive them even of the full quantum of food
corresponding to their weakened capacity of assimilation, and their
descendants will be weakened still more. Inasmuch, now, as no weeding
out of the weaker determinants of the hind leg [or eye] by personal
selection takes place on our hypothesis, inevitably the average
strength of this determinant must slowly but constantly diminish--that
is, the hind leg [or eye] must grow smaller and smaller until it
finally disappears altogether.... Panmixia is the indispensable
precondition of the whole process; for, owing to the fact that persons
with weak determinants are just as capable of life as those with
strong, ... solely by this means is a further weakening effected in the
following generations.”
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