Botany; Geotropism; Plants -- Irritability and movements
downwards; they thus occupy a somewhat different position, relatively
to the centre of the earth and to the branch, from that of the petioles
on the upright branches. With respect to the leaflets, they move at
night towards the apex of the petiole until their midribs stand nearly
parallel to it; and they then lie neatly imbricated one over the other.
Thus half of the upper surface of each leaflet is in close contact with
half of the lower surface of the one next in advance; and all the
leaflets, excepting the basal ones, have the whole of their upper
surfaces and half of their lower surfaces well protected. Those on the
opposite sides of the same petiole do not come into close contact at
night, as occurs with the leaflets of so many Leguminosae but are
separated by an open furrow; nor could they exactly coincide, as they
stand alternately with respect to one another.
The circumnutation of the petiole of a leaf 3/4 of an inch in length,
on an upright branch, was observed during 36h., and is shown in the
preceding diagram (Fig. 136). On the first morning, the leaf fell a
little and then rose until 1 P.M., and this was probably due to its
being now illuminated through a skylight from above; it then
circumnutated on a very small scale round the same spot until about 4
P.M., when the great evening fall commenced. During the latter part of
the night or very early on the next morning the leaf rose again. On the
second day it fell during the morning till 1 P.M., and this no doubt is
its normal habit. From 1 to 4 P.M. it rose in a zigzag line, and soon
afterwards the great evening fall commenced. It thus completed a double
oscillation during the 24 h.
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