Botany; Geotropism; Plants -- Irritability and movements
We then followed a better plan. Holes were bored near the narrow end of
two wooden clips or pincers (Fig. 56), kept closed by brass spiral
springs. Two radicles in damp sand were allowed to grow through these
holes. The
pincers rested on glass-plates to lessen the friction from the sand.
The holes were a little larger (viz..14 inch) and considerably deeper
(viz..6 inch) than in the trials with the sticks; so that a greater
length of a rather thicker radicle exerted a transverse strain. After
13 days they were taken up. The distance of two dots (see the figure)
on the longer ends of the pincers was now carefully measured; the
radicles were then extracted from the holes, and the pincers of course
closed. They were then suspended horizontally in the same manner as
were the bits of sticks, and a weight of 1500 grams (or 3 pounds 4
ounces) was necessary with one of the pincers to open them to the same
extent as had been effected by the transverse growth of the radicle. As
soon as this radicle had slightly opened the pincers, it had grown into
a flattened form and had escaped a little beyond the hole; its diameter
in one direction being 4.2 mm., and at rightangles 3.5 mm. If this
escape and flattening could have been prevented, the radicle would
probably have exerted a greater strain than the 3 pounds 4 ounces. With
the other pincers the radicle escaped still further out of the hole;
and the weight required to open them to the same extent as had been
effected by the radicle, was only 600 grams.
With these facts before us, there seems little difficulty in
understanding how a radicle penetrates the ground. The apex is pointed
and is protected by the root-cap; the terminal growing part is rigid,
and increases in length with a force equal, as far as our observations
can be trusted, to the pressure of at least a quarter of a pound,
probably with a much greater force when prevented from bending to any
side by the surrounding earth. Whilst thus increasing in length it
increases in thickness, pushing away the damp
earth on all sides, with a force of above 8 pounds in one case, of 3
pounds in another case. It was impossible to decide whether the actual
apex exerts, relatively to its diameter, the same transverse strain as
the parts a little higher up; but there seems no reason to doubt that
this would be the case. The growing part therefore does not act like a
nail when hammered into a board, but more like a wedge of wood, which
whilst slowly driven into a crevice continually expands at the same
time by the absorption of water; and a wedge thus acting will split
even a mass of rock.
Public-domain text, read in full here on John Shaqi.
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