The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
then usually of no consequence: indeed the loss is often not rapid enough
to keep the supply from rising to an inconvenient excess. It is chiefly in
respect of that maintenance of heat which is needful during quiescence,
that large organisms have an advantage over small ones in this relatively
diminished loss. Thus these two subsidiary relations between degrees of
growth and amounts of expended force, being in antagonism, we may conclude
that their differential result does not greatly modify the result of the
chief relation.
Comparisons of these deductions with the facts appear in some cases to
verify them and in other cases not to do so. Throughout the vegetal
kingdom, there are no distinct limits to growth except those which death
entails. Passing over a large proportion of plants which never exceed a
comparatively small size, because they wholly or partially die down at the
end of the year, and looking only at trees that annually send forth new
shoots, even when their trunks are hollowed by decay; we may ask--How does
growth happen here to be unlimited? The answer is, that plants are only
accumulators: they are in no very appreciable degree expenders. As they do
not undergo waste there is no reason why their growth should be arrested by
the equilibration of assimilation and waste. Again, among animals there
are sufficient reasons why the correspondence cannot be more than
approximate. Besides the fact above noted, that there are other varying
relations which complicate the chief one. We must bear in mind that the
bodies compared are not truly similar: the proportions of trunk to limbs
and trunk to head, vary considerably. The comparison is still more
seriously vitiated by the inconstant ratio between the constituents of
which the body is composed. In the flesh of adult mammalia, water forms
from 68 to 71 per cent., organic substance from 24 to 28 per cent., and
inorganic substance from 3 to 5 per cent.; whereas in the foetal state, the
water amounts to 87 per cent., and the solid organic constituents to only
11 per cent. Clearly this change from a state in which the force-evolving
matter forms one-tenth of the whole, to a state in which it forms two and a
half tenths, must greatly interfere with the parallelism between the actual
and the theoretical progression. Yet another difficulty may come under
notice. The crocodile is said to grow as long as it lives; and there
appears reason to think that some predaceous fishes, such as the pike, do
the same. That these animals of comparatively high organization have no
definite limits of growth, is, however, an exceptional fact due to the
exceptional non-fulfilment of those conditions which entail limitation.
What kind of life does a crocodile lead? It is a cold-blooded, or almost
cold-blooded, creature; that is, it expends very little for the maintenance
of heat. It is habitually inert: not usually chasing prey but lying in wait
for it; and undergoes considerable exertion only during its occasional
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