The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
In a kindred manner there results a limitation of growth in a land-animal,
which does not exist for an animal living in the water. If, after comparing
the agile movements of a dog with those of a cow, the great weight of which
obviously prevents agility; or if, after observing the swaying flesh of an
elephant as it walks along, we consider what would happen could there be
formed a land-animal equal in mass to the whale (the long Dinosaurs were
not proportionately massive) it needs no argument to show that such a
creature could not stand, much less move about. But in the water the strain
put upon its structures by the weights of its various parts is almost if
not quite taken away. Probably limitation in the quantity of food
obtainable becomes now the chief, if not the sole, restraint.
And here we may note, before leaving the topic, something like a converse
influence which comes into play among creatures inhabiting the water. Up to
the point at which muscles tear from over-strain, larger and smaller
creatures otherwise alike, remain upon a par in respect of the relative
amounts of energy they can evolve. Had they to encounter no resistance from
their medium, the implication would be that neither would have an advantage
over the other in respect of speed. But resistance of the medium comes into
play; and this, other things equal, gives to the larger creature an
advantage. It has been found, experimentally, that the forces to be
overcome by vessels moving through the water, built as they are with
immersed hinder parts which taper as fish taper, are mainly due to what is
called "skin-friction." Now in two fish unlike in size but otherwise
similar skin-friction bears to the energy that can be generated, a smaller
proportion in the larger than in the smaller; and the larger can therefore
acquire a greater velocity. Hence the reason why large fish, such as the
shark, become possible. In a habitat where there is no ambush (save in
exceptional cases like that of the _Lophius_ or Angler) everything depends
on speed; and if, other things equal, a larger fish had no mechanical
advantage over a smaller, a larger fish could not exist--could not catch
the requisite amount of prey.
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