The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
Total insubordination among the centres of development, is shown where the
units or cells, as fast as they are severally formed, part company and lead
independent lives. This, in the vegetal kingdom, habitually occurs among
the _Protophyta_, and in the animal kingdom, among the _Protozoa_. Partial
insubordination is seen in those somewhat advanced organisms, that consist
of units which, though they have not separated, have so little mutual
dependence that the aggregate they form is irregular. Among plants, the
Thallophytes very generally exemplify this mode of development. Lichens,
spreading with flat or corrugated edges in this or that direction as the
conditions determine, have no manifest co-ordination of parts. In the
_Algæ_ the Nostocs and various other forms similarly show us an
unsymmetrical structure. Of _Fungi_ we may say that creeping kinds display
no further dependence of one part on another than is implied by their
cohesion. And even in such better-organized plants as the _Marchantia_, the
general arrangement shows no reference to a directive centre. Among animals
many of the Sponges in their adult forms may be cited as devoid of that
co-ordination implied by symmetry: the units composing them, though they
have some subordination to local centres, have no subordination to a
general centre. To distinguish that kind of development in which the whole
product of a germ coheres in one mass, from that kind of development in
which it does not, Professor Huxley has introduced the words "_continuous_"
and "_discontinuous_;" and these seem the best fitted for the purpose.
Multicentral development, then, is divisible into continuous and
discontinuous.
From central development we pass insensibly to that higher kind of
development for which _axial_ seems the most appropriate name. A tendency
towards this is vaguely manifested almost everywhere. The great majority
even of _Protophyta_ and _Protozoa_ have different longitudinal and
transverse dimensions--have an obscure if not a distinct axial structure.
The originally spheroidal and polyhedral units out of which higher
organisms are mainly built, usually pass into shapes that are subordinated
to lines rather than to points. And in the higher organisms, considered as
wholes, an arrangement of parts in relation to an axis is distinct and
nearly universal. We see it in the superior orders of Thallophytes; and in
all the cormophytic plants. With few exceptions the _Coelenterata_ clearly
exhibit it; it is traceable, though less conspicuously, throughout the
_Mollusca_; and the _Annelida_, _Arthropoda_, and _Vertebrata_ uniformly
show it with perfect definiteness.
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