The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
The bud of any common flowering plant in its earliest stage, consists of a
small hemispherical or sub-conical projection. While it increases most
rapidly at the apex, this presently develops on one side of its base, a
smaller projection of like general shape with itself. Here is the rudiment
of a leaf, which presently spreads more or less round the base of the
central hemisphere or main axis. At the same time that the central
hemisphere rises higher, this lateral prominence, also increasing, gives
rise to subordinate prominences or lobes. These are the rudiments of
stipules, where the leaves are stipulated. Meanwhile, towards the other
side of the main axis and somewhat higher up, another lateral prominence
arising marks the origin of a second leaf. By the time that the first leaf
has produced another pair of lobes, and the second leaf has produced its
primary pair, the central hemisphere, still increasing at its apex,
exhibits the rudiment of a third leaf. Similarly throughout. While the germ
of each succeeding leaf thus arises, the germs of the previous leaves, in
the order of their priority, are changing their rude nodulated shapes into
flattened-out expansions; which slowly put on those sharp outlines they
show when unfolded. Thus from that extremely indefinite figure, a rounded
lump, giving off from time to time lateral lumps, which severally becoming
symmetrically lobed gradually assume specific and involved forms, we pass
little by little to that comparatively complex thing--a leaf-bearing shoot.
Internally, a bud undergoes analogous changes; as witness this
account:--"The general mass of thin-walled parenchymatous cells which
occupies the apical region, and forms the _growing point_ of the shoot, is
covered by a single external layer of similar cells, which increase in
number by the formation of new walls in one direction only, perpendicular
to the surface of the shoot, and thus give rise only to the _epidermis_ or
single layer of cells covering the whole surface of the shoot. Meanwhile
the general mass below grows as a whole, its constituent cells dividing in
all directions. Of the new cells so formed, those removed by these
processes of growth and division from the actual apex, begin, at a greater
or less distance from it, to show signs of the differentiation which will
ultimately lead to the formation of the various tissues enclosed by the
epidermis of the shoot. First the pith, then the vascular bundles, and then
the cortex of the shoot, begin to take on their special characters."
Similarly with secondary structures, as the lateral buds whence leaves
arise. In the, at first, unorganized mass of cells constituting the
rudimentary leaf, there are formed vascular bundles which eventually become
the veins of the leaf; and _pari passu_ with these are formed the other
tissues of the leaf. Nor do we fail to find an essentially parallel set of
changes, when we trace the histories of the individual cells. While the
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account