The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
Under Function in its widest sense, are included both the statical and the
dynamical distributions of force which an organism opposes to the forces
brought to bear on it. In a tree the woody core of trunk and branches, and
in an animal the skeleton, internal or external, may be regarded as
passively resisting the gravity and momentum which tend habitually or
occasionally to derange the requisite relations between the organism and
its environment; and since they resist these forces simply by their
cohesion, their functions may be classed as _statical_. Conversely, the
leaves and sap-vessels in a tree, and those organs which in an animal
similarly carry on nutrition and circulation, as well as those which
generate and direct muscular motion, must be considered as _dynamical_ in
their actions. From another point of view Function is divisible into the
_accumulation of energy_ (latent in food); the _expenditure of energy_
(latent in the tissues and certain matters absorbed by them); and the
_transfer of energy_ (latent in the prepared nutriment or blood) from the
parts which accumulate to the parts which expend. In plants we see little
beyond the first of these: expenditure being comparatively slight, and
transfer required mainly to facilitate accumulation. In animals the
function of _accumulation_ comprehends those processes by which the
materials containing latent energy are taken in, digested, and separated
from other materials; the function of _transfer_ comprehends those
processes by which these materials, and such others as are needful to
liberate the energies they contain, are conveyed throughout the organism;
and the function of _expenditure_ comprehends those processes by which the
energy is liberated from these materials and transformed into properly
co-ordinated motions. Each of these three most general divisions includes
several more special divisions. The accumulation of energy may be separated
into _alimentation_ and _aeration_; of which the first is again separable
into the various acts gone through between prehension of food and the
transformation of part of it into blood. By the transfer of energy is to be
understood what we call _circulation_; if the meaning of circulation be
extended to embrace the duties of both the vascular system and the
lymphatics. Under the head of expenditure of energy come _nervous actions_
and _muscular actions_: though not absolutely co-extensive with expenditure
these are almost so. Lastly, there are the subsidiary functions which do
not properly fall within any of these general functions, but subserve them
by removing the obstacles to their performance: those, namely, of
_excretion_ and _exhalation_, whereby waste products are got rid of. Again,
disregarding their purposes and considering them analytically, the general
physiologist may consider functions in their widest sense as the
correlatives of tissues--the actions of epidermic tissue, cartilaginous
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