The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
Such answer to this question as we may hope to find, must be looked for in
the effects wrought on the organism as a whole by increased function in one
of its parts. For since the discharge of its function by any part is
possible only on condition that those various other functions on which its
own is immediately dependent are also discharged, it follows that excess in
its function presupposes some excess in their functions. Additional work
given to a muscle implies additional work given to the branch arteries
which bring it blood, and additional work, smaller in proportion, to the
arteries from which these branch arteries come. Similarly, the smaller and
larger veins which take away the blood, as well as those structures which
deal with effete products, must have more to do. And yet further, on the
nervous centres which excite the muscle a certain extra duty must fall. But
excess of waste will entail excess of repair, in these parts as well as in
the muscle. The several appliances by which the nutrition and excitation of
an organ are carried on, must also be influenced by this rhythm of action
and reaction; and therefore, after losing more than usual by the
destructive process they must gain more than usual by the constructive
process. But temporarily-increased efficiency in these appliances by which
blood and nervous force are brought to an organ, will cause extra
assimilation in the organ, beyond that required to balance its extra
expenditure. Regarding the functions as constituting a moving equilibrium,
we may say that divergence of any function in the direction of increase,
causes the functions with which it is bound up to diverge in the same
direction; that these, again, cause the functions which they are bound up
with, also to diverge in the same direction; and that these divergences of
the connected functions allow the specially-affected function to be carried
further in this direction than it could otherwise be--further than the
perturbing force could carry it if it had a fixed basis.
It must be admitted that this is but a vague explanation. Among actions so
involved as these, we can scarcely expect to do more than dimly discern a
harmony with first principles. That the facts are to be interpreted in some
such way, may, however, be inferred from the circumstance that an extra
supply of blood continues for some time to be sent to an organ that has
been unusually exercised; and that when unusual exercise is long continued
a permanent increase of vascularity results.
§ 69. Answers to the questions--Why do these adaptive modifications in an
individual animal soon reach a limit? and why, in the descendants of such
animal, similarly conditioned, is this limit very slowly extended?--are to
be found in the same direction as was the answer to the last question. And
here the connexion of cause and consequence is more manifest.
Public-domain text, read in full here on John Shaqi.
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